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Data to the flora of Heves–Borsod Plain I. Distribution of forest, forest steppe and steppe elements
16-65Views:210This paper reports new floristic data of 159 taxa (158 species and one hybrid) from the Heves–Borsod Plain (5 micro-regions between the Mátra and Bükk Mountains and the Tisza River valley). The data were collected between 1999 and 2018 and supplemented with collected specimen (altogether 93 herbarium sheets). This study is the first part of a thematic series that analyzes the distribution patterns of forest, forest steppe and dry grassland ‘steppe’ species, beyond reporting floristic data. The data were derived from 51 flora-mapping quadrats (CEU), based on 5.395 field collected data records. Some indicator species of the three species groups have been assessed in detail (with the addition of distribution maps of 25 species). I have also tested two East-West direction ‘lines’ (the 100-meter isoline and the Csörsz Ditch) for the possible existence of North-South chorological gradients in the area. True forest species (mostly Querco-Fagetea elements) show a dispersed pattern, their representatives can be linked to forest blocks appearing in the landscape. Streams from the adjacented hilly areas (even if they are strongly modified) are very important in the dispersal of the forest species. Their significance and role are decreasing towards the South. For some steppe and forest steppe species (e.g. Brachypodium pinnatum, Campanula bononiensis, Clematis recta, Elymus hispidus, Lychnis viscaria, Ranunculus illyricus, Sanguisorba minor, Stipa spp., Teucrium chamaedrys, Trifolium alpestre, Vinca herbacea) the examined lines indicate a regional area boundary, while for other species (e.g. Phlomis tuberosa, Thalictrum minus) gradient-like distribution differences were not observed. In the distribution of many species an additional southern gradient running along the northern boundary of the former Heves Floodplain was detected. Southwards to this line, the representatives of the selected species are already very sporadic, occurring only in synantropic habitats (e.g. Vincetoxicum hirundinaria).
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The Kámon and Herény source water protection area in the city of Szombathely as a refuge for protected plant species
123–125Views:226The hay meadows and protective forest strips of the Kámon and Herény source water protection areas in the northwestern part of Szombathely have been in good natural condition for decades. By harbouring populations of protected plant species (Orchis morio, Pyrola rotundifolia, Cephalanthera longifolia, Epipactis helleborine and Ophioglossum vulgatum), these areas resemble natural habitats found nearby in the Kőszeg Mountains and the Őrség National Park. The good semi-natural condition can be attributed to the lack of regular mowing in the case of meadows and the lack of disturbance in the case of forest strips. In recent years, however, the area have been exposed to increasing human disturbance (car and motorcycle off-roading), so it may be timely to initiate legal protection. This short case study indicates that properly managed source water protection areas can play an important role in the conservation of semi-natural habitats, rare and protected plant species and the associated fauna.
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Remnants of closed oak woods on loess in the Mezőföld (Pulmonario mollis-Quercetum roboris Kevey 2008)
66–93Views:159The phytosociological characterization of closed oak forests occurring in the loess-covered part of Central Hungary (Mezőföld) is presented. Such forest fragments are found very sporadically in that predominantly treeless region, mostly in semi-dry habitats of north-facing hillsides. This plant community represents an intermediate stage between open steppe woodland (Aceri tatarici-Quercetum pubescentis-roboris) and closed mesic oak-hornbeam forests (Corydali cavae-Carpinetum). Analyses of 20 phytosociological samples using multivariate grouping methods (cluster analysis, principal coordinates analysis) showed a clear difference between these samples and samples of similar communities occurring in the area. In the species composition the characteristic elements of dry and mesic oak forests (Quercetea pubescentis-petraeae, Quercetalia cerridis, Aceri tatarici-Quercion and Fagetalia, respectively) played a major role. On the other hand, dry grassland species (Festuco-Brometea, Festucetalia valesiacae, Festucion rupicolae, etc.) were much less significant than in steppe woodlands. In terms of chorology, European and sub-Mediterranean floristic elements were dominant in the samples, while the proportion of continental elements was substantially smaller than that in steppe woodlands. As a result, our samples were identified with the plant association Pulmonario mollis-Quercetum roboris Kevey 2008, which is classified into the suballiance Polygonato latifolio-Quercenion roboris Kevey 2008 in the phytosociological system.
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Oak-hornbeam and turkey oak forests growing on sand on the foothills of the Pilis Range (Corydali pumilae-Carpinetum, Fraxino orno-Quercetum cerridis)
79–105Views:159We studied the phytosociological characteristics of hornbeam and turkey oak dominated forests growing on sand at the southwestern foot of the Pilis Mountains. Here we present the results of analyses of 10 relevés of each community. The species composition of the hornbeam forest samples is characterized by a relatively high proportion of Fagetalia species. Notable species in the community are Aconitun vulparia, Berberis vulgaris, Epipactis helleborine agg., Galanthus nivalis, Geranium lucidum, Lilium martagon, Omphalodes scorpioides, Piptatherum virescens, Primula veris, Scilla vindobonensis, Sorbus aria agg., S. domestica, Veratrum nigrum, and Waldsteinia geoides. In the turkey oak forests, species characteristic of the class Quercetea pubescentis-petraeae have the highest proportion. Notable and locally typical species occurring in the community are Anthericum ramosum, Digitalis grandiflora, Iris variegata, Piptatherum virescens, Primula veris, Achillea distans, Dictamnus albus, Galanthus nivalis, Lonicera xylosteum, Scilla vindobonensis, Sorbus domestica, Veratrum nigrum, as well as Lunaria annua and Tilia tomentosa that are possibly introduced here. Based on the results of statistical analyses, the two forest communities are best identified with the associations named Corydali pumilae-Carpinetum and Fraxino orno-Quercetum cerridis, both occurring in the Transdanubian Mountain Range.
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Floristical data from loess substrate of Harangod and South-Cserehát (NE Hungary)
105-113Views:77This paper contains the new floristic results of the author from Harangod and SouthCserehát, collected between 2009 and 2013. It presents data on 66 and 48 species respectively. Several new localities of forest-steppe species (e. g. Artemisia pontica, Corydalis cava, Galium odoratum, Lilium martagon, Melica altissima, Thalictrum minus, Viola elatior), meadow species (e. g. Gratiola officinalis, Senecio sarracenicus) and adventiv plants (e. g. Gypsophila perfoliata, Panicum capillare, Typha laxmannii) are published here.
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About the occurrence of Spiraea media Fr. Schm. on the Balaton Uplands
151-154Views:285The presence of Spiraea media on the Balaton Uplands has high plant geographical importance, but the found location has not been known before its recent rediscovery. The Spiraea media was discovered on the Kopasz Hill (Felsőörs) growing in a closed Quercus cerris oak forest (~Fraxino orni-Quercetum cerridis); which is a rather atypical habitat of the species. The appearance of blooming specimens and the rediscovery of the species is probably due to the cut of the neighbouring forest, which has resulted extra irradiation in the area. The 19th century military maps of the landscape show that the former vegetation of the area was characterised by the mosaics of rocky grassland–scrub–forest or rocky scrub rich patches being suitable for the species.
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Taxiphyllum densifolium (Taxiphyllaceae) and other rare bryophytes of the Mecsek valleys (SW Hungary)
3-16Views:88In this paper, we deal with threatened and/or legally protected bryophytes found in the shady, forested valleys of the Mecsek Mountains, which are rare in national and/or local terms. Taxiphyllum densifolium, a rare moss in Europe has stable colonies with sporophytes in two valleys. This study is the first to describe the spore characteristics of this species. The previously considered unique and legally protected Rhynchostegiella teneriffae is nowadays so widespread in the Mecsek that it can no longer be considered a vulnerable species (VU) in Hungary. We present the first population of the legally protected Neckera pennata in Tolna county, and we also report on the recent, unusual occurrence of the data-deficient (DD) Fossombronia wondraczekii. We also list new populations of three other endangered (EN) species, Blindiadelphus recurvatus, Palustriella commutata and Porella arboris-vitae. Several vulnerable (VU) taxa such as Blepharostoma trichophyllum, Porella cordaeana, Sciuro-hypnum flotowianum, Seligeria pusilla, as well as new occurrences of near threatened (NT) mosses such as Microeurhynchium pumilum (with sporophytes), Phaeoceros carolinianus are documented. In addition, the data on Rhynchostegium rotundifolium in Jakab Mountain, which has not been recorded for more than 50 years, is confirmed. The difficulties of differentiating Hygroamblystegium species are briefly discussed.
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Mondró-halom kurgan (Hencida, East Hungary), a refugium of loess grassland vegetation
143-149Views:485Fragments of the natural grassland vegetation are often preserved only in those areas which are inadequate for arable farming. In many cases kurgans hold the last remnants of dry grasslands in lowland areas, like the Great Hungarian Plain. They also have an essential role in preserving cultural and landscape values. Moreover, they harbour several rare plant and animal species. Our aim was to explore the vegetation of the Mondró-halom kurgan (Hencida, East Hungary). Altogether we found 74 vascular plant species in the loess grassland of the kurgan. Several rare species of the Bihari-sík region, such as Inula germanica, Ranunculus illyricus and Rosa gallica were also detected. The steep slopes of the kurgan with various micro-sites and exposures supported a species-rich vegetation. Instead of its small area it harboured several forest steppic species and several steppic flora elements (Continental, Pontic-Mediterranean, Pontic, Pontic-Pannonic and Turanian).
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Epipactis exilis in the Börzsöny Mts (C Hungary)
102-103Views:208Seven specimens of Epipactis exilis has been found in the Börzsöny Mts, in a beech forest along the creek Szén-patak on the 21st August 2020. This strictly protected species is very rare in Hungary, the present observation is its fifth occurrence in the country. It is new to the Börzsöny Mts flora.
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Astragalus vesicarius and other new taxa to the flora of the Vértes Mountains (Hungary, Transdanubian Mts)
195–199Views:186A conservation survey on the condition of steppe and forest-steppe habitats in the Vértes Mts revealed several plant species from the flora of the area that have not been reported before. From a phytogeographical point of view, the most important is Astragalus vesicarius, whose small population in the Vértes fits very well into the native distribution pattern of the species in Hungary. Another important steppe plant is Stipa dasyphylla, which is found in two locations in the Vértes Mts. Besides other taxa of conservation interest (Orobanche teucrii, Phelipanche purpurea, Rosa hungarica), some under-recorded species (Cerastium tenoreanum, Stellaria pallida), and some spreading weeds (Elymus elongatus, Euphorbia lathyris) enrich our knowledge of the flora of the area.
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Ceratocephala testiculata (Crantz) Roth and further data to the flora of the foothills of Bükk Mts.('Bükkalja', NE Hungary)
81-142Views:223This paper reports new floristic data of 367 taxa (365 species and two hybrids) from the foothill region of Bükk Mts. ('Bükkalja', NE-Hungary) based on 5470 field collected data records. The data was collected between 1997–2014 and supplemented with herbaria (altogether 344 herbaria sheets). Data evaluation was based on the comprehensive Bükk Flora of András Vojtkó and further papers. Altogether 45 species were registered as previously unreported from the area, whereas three species had old data (Acer negundo, Elaeagnus angustifolia, Xanthium strumarium). Out of the newly registered species six native species inhabits natural habitats (Carex flacca, Ceratocephala testiculata, Glycerrhiza echinata, Gypsophila paniculata, Kochia laniflora, Marrubium vulgare), whereas five species represents the weed flora (Anthemis ruthenica, Chenopodium ficifolium, Eragrostis pilosa, Senecio vernalis, Xanthium strumarium). The majority of the newly observed species are adventives. Five species were also regarded as recent introduction regionally, although they are native in the country (mostly lowland species: Crypsis alopecuroides, Limonium gmelini subsp. hungaricum, Salsola kali, Scirpoides holoschoenus, Trifolium angulatum). Typical threats observed are landscape and habitat degradation by invasive alien species out of which 11 are newly added. Mass occurrences were detected for the following invasive alien species: Bidens frondosa, Impatiens glandulifera, Phytolacca esculenta, Xanthium italicum. 43 species are also registered, which did not have recently published data from the area. These species include more native elements representing different habitat preferences (dry grasslands, wet meadows, forest and weeds). In case of 65 species the unravelled new occurrences increased significantly, thus contributed to the clarification of their distribution pattern. It was found that the floristic data accumulated on a decade-scale are not sufficient for the analysis of floristic trends, but could serve as a good basis for surveying floristic gradients between the mountainous and lowland areas.
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A new, remarkable population of Epipactis placentina found near Parádsasvár village (Mátra Mts, NE Hungary)
247–248Views:218The present paper surveyes the Hungarian records of Epipactis placentina Bongiorni & Grünanger, which is the rarest Epipactis species in Hungary. Currently, E. placentina is known only at Parádsasvár village in the country. This population (ca. 40 generative individuals) was found in 2019, in an old beech forest, in northern exposition.
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Campylopus flexuosus (Hedw.) Brid. in the Western Mecsek Mts. (South Transdanubia, Hungary)
28-37Views:133During a field survey in 2014, new localities of Campylopus flexuosus, were found in humid acidophilous oak and beech forests in the Western part of the Mecsek Mts. As a new element of the Hungarian bryoflora, C. flexuosus was discovered in 2013 in the Bakony Mts by Peter Erzberger and Csaba Németh. The localities of the sometimes extensive stands are enumerated, the size of the populations is estimated and the associated bryophyte and higher plant species are listed. Some morphological differences between C. flexuosus and the similar moss Dicranum flagellare are described and illustrated. Some interesting results of a revision of D. flagellare specimens in BP are compared with the actual distribution of C. flexuosus and lead to the conclusion that this species can be treated as a potentially expansive moss in humid acidophilous forests of the Mecsek Mts.
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Online distribution atlas of the Hungarian vascular flora (Atlas Florae Hungariae): Homepage structure and prospects of database building
238–252Views:453In December 2018 the online Distribution atlas of vascular plants of Hungary (Atlas Florae Hungariae) was published. The database was built from more than 1 million data records. It mainly contains data from the Hungarian Flora Mapping Programme, but herbarium and literature data were also processed. Maps of species richness illustrate the taxonomic diversity of biogeographical areas and shows the differences of survey quality. After the Flora Mapping Programme ended, between 2016 and 2019, additional data from scientific articles were processed. 25% of new data originated from the article series “Contributions to the Atlas Florae Hungariae” in the Kitaibelia journal, which significantly broadened the distribution of some species. Records of Mesophile forest species (e.g. Allium ursinum, Dryopteris dilatata) increased, as well as records of weed species (e.g. Euphorbia maculata, Senecio vernalis) which can be attributed to their natural spreading. The most important future tasks are to create a reviewed list of taxa and extend data records with an attribute table containing all additional information. Until now the database website is presented only in Hungarian (http://floraatlasz.uni-sopron.hu). Registration is available for anybody, after sending a request via e-mail (atlas.florae.hungariae@uni-sopron.hu). There are different options (excel file by e-mail or online web form) on how to upload floristic data for registered users. Data is only accepted if required information (name of taxa, date, data publisher, settlement, CEU code) is filled out. Each record of uploaded data is revised by the website administrator and taxa specialists. The English version of the webpage will be available in the near future.
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Vulpia bromoides and Luzula multiflora in the Dráva Plain (S Hungary)
234-235Views:1643Vital populations of Vulpia bromoides (L.) S. F. Gray and Luzula multiflora Kirsch. were found between Vajszló and Páprád villages (S Hungary) in 25 May 2022. The populations grew in a mesic forest clearing in the territory of “Bükkhát forest”. These species are new for the flora of the Drava Plain. The habitat is demonstrated by a phytosociological relevé.
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Data to the flora and geobotany of Kisalföld (Lesser Plain) region, NW Hungary
235-253Views:128Lesser Plain (Kisalföld) – divided among three countries (Austria, Hungary and Slovakia) – is the westernmost part of the lowland areas of the Pannonian Basin. Present paper provides a résumé of the authors’ floristic work from the Hungarian part of the area, describing localities, habitats and regional phytogeography of 57 taxa altogether. The localities were listed according to the classification of small geographic regions. The most important results of the study are as follows:
- We recorded 5 species new to the region (Brachypodium rupestre, Bromus ramosus, Hypericum dubium, Lathyrus sphaericus, Tordylium maximum); all of them can be considered as submontane-montane elements.
- We reported several species new to a microregion within the Lesser Plain: montane species in the W-SW part of the Plain (Cirsium rivulare, Dryopteris dilatata, Equisetum telmateia), loess-connected species mainly on the Moson Plateau (Agropyron cristatum, Allium atroviolaceum, Euphorbia salicifolia, Viola ambigua); and occurrences of species on sandy soils near the Lake Fertő (Draba nemorosa, Stipa pennata).
- We explained the known localities of several salt tolerant species (e.g. Carex divisa, Crypsis alopecuroides, Juncus maritimus, Limonium gmelinii, Suaeda pannonica) from the Lake Fertő and its surroundings.
- We assessed the role of the forest management and nature conservational actions in the distribution of some important tree species (Acer tataricum, Betula pubescens, Fraxinus ornus, Quercus petraea, Salix pentandra).
- We discussed the regional nature conservational status of several protected and/or threatened species (e.g. Allium carinatum, Allium suaveolens, Carex strigosa, Hottonia palustris, Potamogeton coloratus, Stellaria palustris) in the Lesser Plain.
The activities of the authors in the surroundings of Győr were supported by monitoring results gained during the Hungarian Little Plain project (LIFE08 NAT/H/000289).
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Habitats and protected plants of Cuha-valley (Bakony Mts, W-Hungary)
80-88Views:66Floristic data were collected and habitat mapping was performed between 2009 and 2011 in the valley of Cuha-creek. During the field work occurrences of 423 vascular plant species (including 31 protected taxa) were recorded. Potentilla indica represents a previously unknown alien taxa of the study area. The dominant vegetation type of the Cuha-valley is beech forest. Further characteristic, important habitats are scree, ravine and hilltop forests. On the southern part of the valley – in close proximity of the Cuha-creek – expansion of several alien plants (Impatiens glandulifera, Fallopia spp., Heracleum mantegazzianum) were observed.
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White poplar riparian forests on the Csepel-sziget, Middle-Hungary (Senecioni sarracenici-Populetum albae Kevey in Borhidi & Kevey 1996)
57–78Views:189In this paper the white poplar riparian forests (Senecioni sarracenici-Populetum albae) growing on the Csepel-sziget and its vicinity are described and characterized based on 25 phytosociological relevés. These communities grow on loose fluvial sand and raw alluvial soils on the elevated parts of the lower river floodplain. They can readily be distinguished from willow gallery forests (Leucojo aestivi-Salicetum albae) which have no shrub layers and grow in habitats 1–1.5 m below the level of poplar forests on rather heavy and muddy soils. They differ also from the oak-ash-elm forests (Scillo vindobonensis-Ulmetum) growing in the upper floodplain. Certain – partly submontane – plants that are rare or completely absent in other parts of the Great Hungarian Plains may also occur in them, such as Anemone ranunculoides, Carex remota, Clematis recta, Crataegus × degenii, Crataegus nigra, Epipactis helleborine, Equisetum hyemale, Galanthus nivalis, Lathraea squamaria, Leucojum aestivum, Paris quadrifolia, Scilla vindobonensis, Vitis sylvestris. This association is classified in the sub-alliance Populenion nigro-albae Kevey 2008 in the syntaxonomical system.
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Data to the distribution and nature conservation of Sorbus bakonyensis
229-233Views:1296Sorbus bakonyensis (Jáv.) Jáv. (syn. S. majeri Barabits) is an apomictic species with a very narrow distribution range in the Bakony Mts (Hungary). Its previous occurrence data were published only from Kopasz Mt. (~Kápolna Hill, Csordás valley) above the village Márkó. It was possible to slightly expand the known area of the species to the East and to the West (Kis-Bükk Mt) of Kopasz Mt. with a detailed mapping. Due to the number of known individuals (six mature trees and a variable number of seedlings) and its small distribution range (~2 km2) the species is critically endangered. Regarding its small population, the high number of big game (wild boar, red deer) and the more and more frequent drought periods are significant risk factors. In situ preservation of the species is still a realistic goal, but ex situ propagation in tree nurseries and reintroduction to the suitable habitats of the mapped area (clearings, forest edges) are also justified.
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A new record of the threatened Anacamptodon splachnoides (Froel. ex Brid.) Brid. from the Bükk Mts. (NE Hungary)
202-205Views:140A new locality of the moss species Anacamptodon splachnoides (Froel. ex Brid.) Brid. was discovered in a Turkey oak-dominated forest stand in the Bükk Mts. (near Varbó village, North Hungarian Montains). A small population of this threatened bryophyte was found in a wet root-hole of a living Quercus cerris L. tree.
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Temperature and soil moisture regimes of the forested and non-forested dolines of the Bükk Mountains based on ecological indicator values
331-338Views:98In the study we aimed to investigate whether there are differences in the ecological conditions of the forested and non-forested dolines of the Bükk Mountains (northern Hungary). Transects for sampling the herb layer were established across three forested and three non-forested dolines in a north to south direction, traversing the deepest point of the dolines. Presence-absence data of each vascular herb and tree sapling were recorded in the plots. Borhidi’s indicator values for temperature (TB) and soil moisture (WB) were used to compare the ecological conditions along the slopes. Our results showed that there were remarkable differences between the temperature and moisture regimes of the forested and non-forested dolines. Both the temperature and moisture gradients were more pronounced along the slopes of the non-forested dolines than along the forested ones. These are due to the fact that forest cover together with the features of the regional climate of the area has a considerable mitigating effect both on the air temperature and soil moisture regimes of dolines.
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New occurrences of Ranunculus strigulosus in East Hungary
3–8Views:330The exact distribution of Ranunculus strigulosus Schur is poorly known in Hungary. Current occurrence data of the species mainly from Békés county and only one archive data from the Nyírség region, from the 1920s were known. This study presents eleven newly discovered occurrences in East Hungary. Populations were found in wet meadows, forest glades and poplar plantations. Nine of these populations are located at the southeastern edge of the Nyírség region and most of them are quite small. Three larger population were found in Nyírábrány, Álmosd and the Tócó Valley between Debrecen and Hajdúböszörmény. One of the newly discovered populations provides a new contribution to the flora of Bátorliget Marsh and Fen-wood NCA. Some morphological characters (hairs on the stem and the leaves, shape of the leaves developed before and after the mowing) are also discussed.
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White willow riparian forests along the upper Tisza River, Hungary
78–100Views:111Forests of the Tisza floodplain in the northwestern part of Hungary include white willow riparian forests (Leucojo aestivi-Salicetum albae) that are phytosociologically little studied. This study summarizes the characteristics of this community based on 25 phytosociological relevés. These forests grow mostly on mud and raw alluvial soil in the low-lying parts of the lower terraces of the floodplain. They are easily separated by their species composition and underdeveloped shrub layer from white poplar gallery forests (Senecioni sarracenici-Populetum albae)., which grow on higher lying ground with rather light sandy soil and typically possess a pronounced shrub layer. The understory of their stands may often host plants that are rare or completely absent in other parts of the country, such as Cardamine amara, Cardamine flexuosa, Cardaminopsis arenosa, Carex pseudocyperus, Carex remota, Chrysosplenium alternifolium, Leucanthemella serotina, Leucojum aestivum, Oenanthe banatica, Scrophularia scopolii, Telekia speciosa, Vitis sylvestris. This community is classified in the „Salicenion albae-fragilis Kevey 2008” suballiance.
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Contributions to the flora of Tolna and Baranya counties IV.
17-33Views:131The authors present the most important floristic data of Tolna and Baranya counties, collected together or separately in the last 15 years, or observed by other data providers. In total, occurrence data of 140 taxa are reported. Most of them are protected species in Hungary, but the list also includes rarer weeds (Adonis flammea, Althaea cannabina, Aphanes arvensis, Bunias orientalis, Legousia speculum-veneris, Vicia lutea) and adventive species (Helminthia echioides, Oxybaphus nyctagineus, Rubia tinctorum). Several species of phytogeographical importance were also found in the forest-steppe vegetation of the loess areas studied. Occurrence of Linaria biebersteinii should be highlighted. L. biebersteinii is a very rare species in the Transdanubian region, and is no longer found in Europe west of this region. The occurrence of Himantoglossum adriaticum is a new contribution to the flora of the southern Transdanubian region.
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New data on the Hungarian adventive flora: Acer opalus Mill. subsp. obtusatum (Waldst. et Kit. ex Willd.) Gams 1925 in Hungary
229-238Views:58Acer opalus Miller 1768 because of its complex taxonomic status can be interpreted as a collective species which contains several species and subspecies depending on different interpretations. An important adventive occurrence from this collective species was found in a forest that belongs to the Tétényi plateau (Budapest, 11th district) in July of 2009. It is proved to be Acer opalus Mill. subsp. obtusatum (Waldst. et Kit. ex Wild.) Gams. The natural range of the subspecies covers the eastern side of the central Mediterranean Sea region and western Balkan Peninsula. It is a submediterranean-mediterranean, thermophile, shade-tolerant taxon but often appears together lightdemanding species too. In its native range it occurs in beech, pine, sessile oak forests and in different types of dry oak forests as admixed tree species. The subspecies was found in a turkey oak – sessile oak community in Hungary. Several old, seed growing individuals live on the spot which have had continuous seed production. Therefore from the old trees to the young seedlings different age categories are represented.