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General principles in variety-association for intensive plantations of pomeceous fruits
96-101.Views:1618Under conditions of Hungary, more than 400 varieties of apple, pear and quince varieties have been observed for time of blooming and fertility relations in order to check the possibility of their use for intense plantations in different combinations with polliniser varieties. Low (below 3%) rate of self-fertility occurred at 65% of apple varieties. That partial self-fertility, however, is far from being sufficient to produce acceptable yield, thus allogamous pollination is absolutely necessary. The same is true for the rear and quince varieties grown in Hungary, too. The normal development requires the presence of viable seeds in the fruit set, most in quince, therefore, association of the right varieties is most important in that species. Apple and pear varieties are assigned according to their blooming time to 4, quince varieties to 3 groups. The yield of all three pomaceous species declines with the growing distance from the potential pollen source. In the intense plantations, the critical (maximum) distance to be observed is 20 m for apple and 15 m for pear and quince. In combining the placement of varieties, also the principles of a variety-specific cultivation are to be considered carefully. The double objectives are satisfied most by the system of Maluspollinisers developed for intense plantations.
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New sweet cherry cultivars in intensive plantings
13-16.Views:965The study took place in the largest sweet cherry plantation in West Hungary. The purpose has been the identification of those varieties, which will be suitable for intense cultivation, early fruiting and excellent fruit quality, moreover, the selection of the optimal phytotechnical procedures. At the same time, scion-rootstock combinations have been tested also from the point of view of growing intensity and fruiting in high-density plantation. The dense planting is induced to start fruiting early and yield regularly by special methods.Yielding was stimulated by maintaining the balance of vegetative-generative growth by binding the shoots, by summer pruning, by cuts on the trunk and root pruning. Best experiences have been found in yield and quality in the following varieties: Canada Giant, Carmen, Firm Red, Giant Red, Katalin, Kordia, Regina. Dense planting has been feasible also on vigorous rootstock, like P. mahaleb. Dwarfing rootstocks like P-HL-A, Gisela 6, accelerate the formation of flower buds and yielding earlier with fruits of adequate size. ‘Firm Red’ and ‘Giant Red’ excelled with their large fruit (>27 mm diameter) in all combinations, thus being promising under Hungarian conditions.
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Fruit drop: I. Specific characteristics and varietal properties of fruit drop
59-67.Views:15230The basic conditions of fruit set (synchronic bloom, transfer of pollen, etc.) still do decide definitely the fate of the flower in spite of the best weather conditions. Beyond a set quantity of fruits, the tree is unable to bring up larger load. A system of autoregulation works in the background and causes the drop of a fraction of fruits in spite of the accomplished fertilisation and the equality of physiological precedents. This study discuss this physiological process based on the international specific literature. The further development of fruits maintained on the tree depends mainly on the growing conditions (e.g. water, supply of nutrients, weather adversities, pruning, fruit thinning, biotic damages, etc.), which may cause on their own turn fruit drop especially at the time of approaching maturity.
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Self-fertility studies of some sweet cherry (Prunus avium L.) cultivars and selections
21-26.Views:273Fruit set of two sweet cherry cultivars (Alex' and 'Stella' ) and four Hungarian selections have been studied over two years following open pollination, unassisted self-pollination (autogamy) and artificial self-pollination (geitonogamy). Among accessions 'IV-6/240' proved to be self-sterile, while the other five revealed to be self-fertile. Significant differences have been found in fruit set among years and among pollination treatments but not among self-fertile accessions. Fruit set following unassisted self-pollination was significantly lower than of other pollination treatments. Thus pollen transfer is essential for profitable yield in sweet cherry growing.
There was no significant relationship in the fruit set of open- and self-pollination.
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Preliminary results of renewal pruning an 18 years old sour cherry trees
41-42.Views:311Sour cherry is a light demanding fruit species. As most of the crop is developing from buds on one year old shoots and 2-yearold wood, the formation of bare wood can be seen apparently in a badly maintained canopy. The formation of „whip shoots” indicates the degree of bare wood. Unfortunately in many gardens and commercial orchards sour cherry trees resemble willow trees. Owners often decide to grub old orchards, although the useful cropping lifespan of the trees planted in a correct spacing can be 20-30 years on a move vigorous rootstock. Therefore renewal pruning can contribute to the full renewal of the orchard, but the length of its effect is depending on the growth characteristics and renewal capabilities of the cultivar. The growth characteristics of the cultivars are different. The regeneration capabilities of different aged wood are also different. These differences emphasize the need for cultivar specifi c pruning. Knowing the reactions to renewal pruning, we can ensure regular high crops with excellent quality by applying cultivar specifi c rotation pruning.
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Impact of sodium-selenate on the growth of radish (Raphanus sativus L.) seedlings in vitro
113-115.Views:547Selenium (Se) is an essential trace element for animals, microorganisms and some other Eukaryotes. It has become increasingly evident that Se plays a significant role in reducing the incidence of lung, colorectal and prostate cancer in humans. Although it is well known that some species among higher plants are able to accumulate selenium in their tissues, but others are not able to do so, and there is evidence that selenium is needed for the growth of algae, meanwhile the question of essentiality of Se in vascular plants is unresolved. We aimed to study the in vitro growing and to characterise some physiological properties in radish (Raphanus sativus L.) seedlings treated with 0 to 200 mg/1 sodium-selenate. The results showed that lower (2 mg/1) concentration sodium-selenate increased the biomass as well as the total antioxidant capacity of seedlings. The seedling's selenium content showed linear correlation with the sodium-selenate content of the medium.
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Floral biological investigations of apple cultivars in relation to fire blight
9-14.Views:371Floral activity was studied in two apple cultivars: an Erwinia-tolerant (Treedorn') and a sensitive one (`Sampion'). Since more types of protogyny occur in apples, the period of stigma activity is different. Papillae of the exposed stigma in flowers of 'Freedom' function longer (usually more than a week) than in the delayed homogamous `Sampion'. Despite of this, cv. 'Freedom' is tolerant to Erwinia amylovora (Burr.) Winslow et al., suggesting no relationship between the floral biological type (including the exposure and longevity of stigma) and the infection by E. amylovora. According to SEM micrographs, nectary stomata in `Freedom' are already open in the flower bud, where nectar secretion starts and continues until the senescence of the stigma. However, the long period of nectar secretion does not create optimal conditions for bacterial growth, since nectar production is scant in the flowers of 'Freedom'. The surface of the nectary, its nectar-retaining capacity, and the amount and concentration of nectar may influence the susceptibility of apple cultivars. It is manifested well by the smooth nectary surface with nectary stomata rising slightly above the epidermis in flowers of cv. 'Freedom', contrasting the wrinkled, striate nectary surface with slightly sunken stomata in the flowers of 'Sampion'.
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Evaluation of precision farming with regard to horticulture
105-110.Views:232The global positioning system was made available for public use, which made it possible' to apply a new management tool in agriculture. Precision farming gives much more information on plant-growing than former methods, which makes it possible to use technologies more suitable for micro-sites. It is supposed that more profitable production can be realised with its aid and the strain on the environment can be reduced, not to mention other economic advantages.
The study makes economic conclusions about the method more and more widely used in plant production taking the yield mapping of Józsefmajor Experimental Farm as a basis and starts ideas about its possible application in arable land olericulture, viticulture and fruit production.
These issues are important because precision farming has existed in glass-culture for a long time from another perspective, which should be spread to open ground horticultural enterprises that are labour and asset intensive, qualitative farming forms with great plantation value.
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Legal harmonization of the Hungarian horticulture conditioned by the accession
23-28.Views:313In the EU-horticulture is treated with special care. As far as its regulation is concerned, different orders are in force to single branches. The regulation of the branch of medicinal plants is related chiefly to the processing, because the overwhelming part of the basic material is to be imported. More preoccupation falls to the ornamental plant branch, although both the production and the turnover are controlled by the market. Irrespective of this fact, the quality standards are high. Also the production and the sale relations are carefully circumscribed, with special regard to the processing procedures. In the control of the quality an important role devolves on the producers' organizations, likely in our country on the so-called TESZ-es (Cooperatives for production and sale). In this regard the domestic prescriptions took much over of the EU practices.
The structure of sales in vegetable and fruit branches is continously changing, the demands of the consumers are more and more satisfied by the super- and hypermarkets. Their marketing organizations offer almost the half of the total of quantity commodities. In consent with the processing industry they raise quite severe requirements to producers. These requirements are to be taken into account also among our circumstances. There is a similar situation valid in the viticulture and winery. Within these branches we are able to compete on good chances of adapting our regulations to those of the EU. In this area there falls also responsibility to our vine-growing communities. Particulary a watch must be kept over the reputation of our renown quality wines and the results already achieved must be protected.
The competent ministries — at first the Ministry of Agriculture and Rural Development — has already shown great flexibility to the desires in regard to the expectable legal harmonization. The legislative, registrative and controlling activities are to be continued in this mentality.
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Effect of the environmentally friendly production systems on the vegetative characteristics of apple cultivars
11-14.Views:578In this study effect of the environmentally friendly production systems on the brunch cross sectional area of apple trees trained to free spindle canopy are presented based on the data of 39 cultivars. According to our results branch thickness of the apple cultivars located in the lower, middle and upper region of the tree are more homogenous in the integrated production system compared to the organic one. Therefore the severity of the annual maintain pruning must be necessarily more moderate in the integrated growing system, than in the organic production system.
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Variety specific integrated fruit production development in order to optimize inner content values
27-31.Views:418In this paper we introduce our results of three years (2007-2009) investigatio ns carried out in the framework of "Research and development in foodstaff chain" - project of Regional University Knowledge Center by the members of Corvi nus U niversity of Budapest - Department of Pomology and. Research Institute for Fru it growing and Ornamentals. Budapest-Erd. The main objectives of the project were the followings: submi ssion of sour cherry candidate suitable for industrial process for state approval; determination of physica l parameter and inner content value changes of sour cherry varieties duri ng ripening; evaluation of health care attributes of sour cherry fruit; ripen ing process description by the colour and the force required to pick fruits of sour cherry. We stated that the optimal begi nning and period of the harvest can be determined wi th the fruit removal force. In this period the fruit growth stops. ju iciness rat io does not change. refractions are approximately equal.acidic content turns to decreasing trend.However the proportion of anthocyan in and polypheno l can still increase. Fruits harvested in this period fulfil a wide range of industrial process opportunities. 'Erdi jubileum ·and candidate IV-3/48 according to their inner content values arc suitable for high quality products (containing real fruit material in high proportion).
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Fruit drop: II. Biological background of flower and fruit drop
103-108.Views:972The most important components of fruit drop are: the rootstock, the combination of polliniser varieties, the conditions depending of nutrition, the extent and timing of the administration of fertilisers, the moments of water stress and the timing of agrotechnical interventions. Further adversities may appear as flushes of heat and drought, the rainy spring weather during the blooming period as well as the excessive hot, cool or windy weather impairing pollination, moreover, the appearance of diseases and pests all influence the fate of flowers of growing and become ripe fruits. As generally maintained, dry springs are causing severe fruit drop.
In analysing the endogenous and environmental causes of drop of the generative organs (flowers and fruits), the model of leaf abscission has been used, as a study of the excised, well defined abscission zone (AZ) seemed to be an adequate approach to the question. Comparing the effects active in the abscission of fruit with those of the excised leaf stem differences are observed as well as analogies between the anatomy and the accumulation of ethylene in the respective abscission tissues.
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Hungaricum as a quality of fruits and fruit products
71-81.Views:1091The territory of the Hungarian state is largely suitable for the purpose of growing fruits of the temperate zone species. During the next decennia, the annual volume of Hungarian fruit production is expected to be around 1.1-1.3 million tons, from which some 15% is considered to be a produce of Hungary or "Hungaricum" (90 thousand tons of sour cherry, 50 thousand tons of apricot, 20 thousand tons of raspberry, 10 thousand tons of walnut). These fruits symbolise the country's special quality, which are worth to catch the interest the foreign consumers.
The category of Hungaricum involves almost exclusively varieties of Hungarian origin as sour cherries, apricots, raspberries and walnuts, and they are representing outstanding qualities on the international markets.
As for the fruit products the fruit brandies are eligible to be "Hungaricum" and are called exclusively "Pálinka". The Pálinka, provided to be distinguished with a geographic mark and will be competitive on the world market. Smaller quantities, though significant produce is represented by the deep frozen raspberry.
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Volatile constituents of Nepeta cataria L., N. glechoma Benth. and N. parviflora M. Bieb. from Hungary
47-50.Views:238In the temperate zone live about 150 species of the Nepeta genus. Our investigations covered the examinations of the volatile oil containing species of the genus endemic in Hungary, Nepeta cataria and Nepeta parviflora. Latter is a relict of the ancient steppe-flora and endemic in Hungary as well. Phytochemical examination of the volatile oil containing plant material has also been carried out. Catnip growing in the Botanical Garden of PTE Department of Botany contained 0,67% volatile oil in May and 0,14% in November. Chemical character of the volatile oils were measured by gas chromatography/mass spectrometry and citronellol, citral-A, citral-B and geraniol components were identified. The composition of the oil of November samples shifted towards citronellol (65%). In both samples insecticide and repellent activity bearing compounds (+)-cis-p-menthane-3,8-diol, and (—)-trans-p-menthane-3,8-diol in 2-2.5 and 4-4.5% amount have been found. The catnip sample deriving from Germany contained a small amount of anetol, citronellol, neral, geraniol and geranial (6-13%), and possibly two isomers of nepetalactone in 23-31%.
The Nepeta parviflora endemic in the Nagyvolgy valley near Nagykaracsony consisted of the same compounds in the investigated years (1998-2000). Its limonene, methyl chavicol, b-cariophyllene, b-selinene, b-cubebene, davanone, germacrene-D constituents have been identified. In the year 2000 different GC % of these compounds were detected in the different organs of the plants.
The closely related species Nepeta cataria var. citriodora contained 83% citral, and the N. glechoma (= Glechoma hederacea) contained 41% a-cubebene, 20% patchoulenol, 7,7% spathulenol respectively. These compounds were identified by gas chromatography and gas chromatography / mass spectrometry.
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Evaluation of hot pepper (Capsicum annum L.) varieties for green pod yield and quality under rain fed production at Teppi, South Western Ethiopia
45-49.Views:383The trials were designed with three replications in a randomized complete block design in order to evaluate the phenological, growth and yield potential of hot pepper varieties such as ‘Mareko fana’, ‘Melka Zala’, ‘Melka Awaze’, ‘Melka Shote’ and local check. Significant difference was observed between growing year and used varieties. The varieties also performed significantly different (p<0.05) for most of the considered traits in the study. The result revealed that varieties ‘Melka Zala’, ‘Mareko fana’ and ‘Melka Awaze’ were scored highest green pod yield of 8.39, 8.71 and 11.39 ton per hectare, respectively. However, ‘Mareko fana’ variety was susceptible to disease attack as compared to other varieties. Therefore, promoting both ‘Melka Awaze’ and ‘Melka Zala’ varieties for widespread production for Teppi and the areas with similar agro-ecological conditions could contribute to boost the productivity of hot pepper. ‘Mareko fana’ could also be used for dry pod purpose due to its attractive color.
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Evaluation of the vegetative and generative performance of new apple cultivars in the Nyírség region
39-44.Views:451In this paper we examine apple varieties, which are planted expansively in the development apple orchards of the Western- European countries, but about its production characteristics we don’t have or just few practical experiences in Hungary. These varieties can be described with regular and high yields, aesthetic and alluring appearance in their origin places, what ensure profitable producing for the growers. Under the Hungarian climate which tends to be often extreme (hard cold winters, late spring frosts, summer drought, heat stress) these varieties may behave differently, so before the substantial planting it is expedient to carry out variety evaluations to avoid the later failures. The place of our experiments is located in the Nyírség representing adequately the ecological conditions of the surrounding areas, so the gained results can be adapted easily in the biggest apple production site of Hungary. Regarding the vegetative and the generative parameters of the 14 examined varieties (’Gala Venus Fengal’, ’Gala Decarli-Fendeca’, ’Galaval’, ’Jugala’, ’Gala Schnitzer (S) Schniga’, ’Red Cap Valtod (S)’, ’Early Red One’, ’Jeromine’, ’Crimson Crisp (Co-op 39)’, ’Red Topaz’, ’Wilton’s Red Jonaprince’, ’Red Idared’, ’Fuji September Wonder’, club cultivar) we found significant differences. The ratio of the thickness of trunk and the main branches showed that the trees have an optimal canopy structure in accordance with the Zahn principles, so they are in harmonious growing balance. The ‘Gala’ sport trained to slender spindle produced the highest yields, while the ‘Fuji’ and the ‘Red Delicious’ sport trained to super spindle can be described with the lowest values. All the cultivars reached the eating quality (65-70 mm), from which the ’Red Idared’, the ’Red Jonaprince’ and the ’Red Cap’ produced outstanding fruit size (81-85 mm).
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The use of rootstocks for European (Prunus domestica) and for Japanese (Prunus salicina) plums (review)
7-13.Views:1090The worldwide tendency to increase the intensity of fruit growing technologies prefers generally for every fruit species rootstocks with week or mediocre vigour. From this viewpoint, the use of rootstocks for plums are rather unilateral in Hungary, where 95–99% of plum plantations are grafted on mirobalan seedlings (P. cerasifera v. mirobalana). The score of plum rootstocks abroad is much more diversified. The present study summarises the respective knowledge referring to the literature available.
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Comparative organoleptic examination of apple varieties developed by different environmentally safe technologies
69-70.Views:388Ecological or biological growing techniques are generally recognised as actual trends in agricultural production. The objective of our experiments is a comparison of apple fruits grown alternatively by integrated versus ecological technology. In a collection of varieties, the possibility of the experiment was feasible. At the same time, a comparison of the popularity of existing varieties was also actual. The organoleptic consumer’s tests rated the varieties, and the comparison with instrumentally raised “objective” data could be compared with the subjective ratings, which did not coincide with each other. This phenomenon was revealed in the rating of acid-sugar harmony, where the internationally used quality index of Thiault produced a different range of the samples then the consumer’s test. The same contradiction was observed between ratings of firmness and the data raised by penetrometer.
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Molecular identification of old Hungarian apple varieties
37-42.Views:751Altogether 40, mainly old Hungarian apple varieties were screened with six previously described microsatellite markers. A total of 71 polymorphic alleles were detected (average 11.8 alleles/locus) and the heterozygosity of markers averaged very high (0.8). The genetic variability among the genotypes proved to be so remarkable that as few as three markers from the applied six were enough to distinguish between the 40 varieties. This was also confirmed by the cumulative probability of obtaining identical allele patterns for two randomly chosen apple genotypes for all loci, which value was quite low: 2.53 x 10-5. The molecular identification of these genetically very different old apple genotypes could be very useful in future breeding programs.
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The effect of rootstock on the tree size of apricot cultivars
57-58.Views:245The apricot is propagated on several kinds of rootstocks in Hungary. The main aspects of selecting rootstocks are as follows: adaptability to environmental circumstances, primarily soil conditions, ensuring the tree size that complies with the cultivation method, and compatibility with the grafted cultivar. At advanced, intensive orchards rootstocks ensuring smaller tree size are privileged. For the establishment of the appropriate cultivation system, it is important to be aware of the expected growing vigour and tree size of certain cultivarrootstock combinations when the orchard is designed. In the course of our experiment the size of 15-year-old trees of 4 apricot cultivars were examined on several rootstocks at an orchard in Siófok. On the basis of the data measured for each cultivar-rootstock combination, it can be stated that trees on wild apricot (P. armeniaca) rootstocks are the largest in size. Trees on prune (P. domestica) rootstock have 10-15% smaller crown volume than the previous combination. Trees on bullace (P. insititia) rootstock have the smallest tree size and their crown volume is 30-50% smaller than that of the trees on P. armeniaca rootstock. Thus, the application of prune and bullace rootstocks is beneficial at intensive apricot orchards as the size of trees can be reduced by their usage. However, their compatibility with the cultivars and their adaptability to the ecological conditions of the production site have to be tested before applying them widely. In the course of our research incompatibility was not experienced for any of the cultivar-rootstock combinations examined. Nevertheless, the drought tolerance of the rootstocks examined showed significant differences. Trees on P. domestica or P. insititia rootstock requires more water than those on P. armeniaca rootstock, therefore, they have to be irrigated.
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Studies on the insect pollination of fruit tree species and on closely related topics in Hungary: a bibliography of research publications
157-161.Views:330In the past decades fruit production has become more and more intensive and so the interest towards the bee pollination of fruit trees has increased in Hungary. Thus insect pollination research has become more intensive in the eighties and in the nineties. Recently, high density fruit orchards are planted of dwarf trees at more and more farms in Hungary and these need highly controlled growing practices involving insect pollination, too. This is a new challenge of insect pollination research in Hungary. So it seems to be worth to compile a list on research publications having been appeared so far to create a basis to the ongoing new research. Besides papers strictly on the insect pollination of fruit trees publications on some closely related topics, first of all on the nectar production and on other flower characteristics of fruit tree cultivars that directly influence bee activity have also been included on the list.
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Disease warning models for brown rot fungi of fruit crops
19-22.Views:606In this review, disease warning models for brown rot fungi, including Monilinia fructigena, M. laxa and M. fructicola, were summarized. Few studies have been made to relate epidemiology and disease warning in brown rot infection caused by M. fructicola and M. laxa in order to predict infections or develop decision support models for fungicide applications during the growing season. More recently a disease warning model and a decision support system were also performed for M. fructigena for organic apple orchards. This review gives an overview on some details of the above disease warning models and decision support system.
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Effects of excessive weather on the micro-climate of apple plantations under the hail protection nets
81-85.Views:26154The general utilisation of hail nets is spreading all over the world and in Hungary too. Hail nets are at present the only tool, which
is able to prevent heavy hail damage in agricultural crops. Besides its numerous advantages, there are also deleterious side effects, e.g. it
changes the microclimate of the crop stand. Our study aimed to observe the differences, which characterise the physical status of crop stands
under the protection of hail nets compared with the stands in open air under various meteorological conditions: high or low temperature, calm
or windy atmosphere and their possible combinations. The results indicated that on hot and calm days the difference may attain 3–4° C. On
cool summer days, the difference was only 1.5 ºC. The hail net influences the relative humidity of the air, which means on ho summer days
often 7-8% differences. Conditions of radiation may also differ conspicuously. Measurements prove the reduced global radiation on sunny
summer days to 70%. This impairs as a rule the development of fruits reducing their quality and also its quantitative traits. Sunburn is, on the
other hand, significantly prevented by shadow. -
Organoleptic evaluation of pear varieties grown at different sites
33-37.Views:973It was stated as a summary that a system of testing pear varieties on scale of 100 degrees, weighing 11 properties, gives a comprehensive information on qualities of pear varieties. The points of the form to be filled out, distinguishes the appreciation of accessory flavours and tastes from the main properties. Under the title of taste, sweet and acid taste are evaluated separately. The presently used system is a step forward as the composed characters do not influence the judgement of each other. In the representation of results, the 100 degree system was more efficacious than the usual 5-point system because there was no need of using fractions, and the graphic profile is more expressive, so the commodity is better visualised. The assembly of the jury was divided into two groups as below and above 35 year old persons. The two groups judged differently the acidity, sweetness and the thickness of the fruit peel. It was suggested that the choice of the produce should consider the differences in taste and give up the idea of being equally favourable for both groups. The samples of different growing sites revealed that the best quality is produced at Nagykanizsa, and this capacity would be able to supply the market also in the large super-markets and chain of department stores in Hungary.
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Reproductive biology of Duranta repens L. (Verbenaceae) in relation to its environment
51-55.Views:416Impact of environmental changes (temperature and RH) on reproductive biology in Duranta repens (Verbenaceae) growing at ten different sites of Agra was studied. It flowers throughout the year, with optimum flowering in September. The flowers are arranged in loose clusters on terminal or axillary racemes. They are either blue or lavender in colour, hermaphrodite, actinomorphic and complete. The plants exhibit floral polymorphism (increase and decrease in number of petals and stamens) and considerable variation in extent of pollen fertility, floral density, insect pollinators and fruit-set percentage. The changes in temperature and relative humidity during the entire flowering period, was found associated with the variation in floral structure, pollen fertility and fruit-set percentage. Based on the percentage of fruit-set during different seasons of a year, there were three distinct periods, namely maximum, moderate and minimum periods. The present paper deals with the comparative view of reproductive biology of this ornamental plant in these periods. During the months of August—November when temperature ranges between 13.7-36.6 °C and RH between 79-89% the plant exhibits maximum fruit-set percentage (68-85%). This was associated with maximum flowering, increase in floral size, and increase in visitation rates of pollinators and higher degree of pollen fertility. On the other hand, with temperature reaching to the maximum (15.1-41.5 °C) and reduction in RH (14.1-41.3%), the percentage of fruit-set was reduced to the minimum (21-30%). During this period, number of flowers/plant, floral size, pollen fertility, visitation rates of pollinators were reduced to the minimum. During this period floral polymorphism was also 'recorded.
