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Toxicity of fungicides to honey bees (Hymenoptera: Apidae) and their effects on bee foraging behavior, pollen viability and fruit set on blooming apples and pears
96-98.Views:311Fungicides fosetyl-AL, triadimefon, dodine, mycobutanil and fenarimol were tested for honey bee (Apis mellifera L.) mortality and effect on bee foraging, pollen viability and fruit set in blooming apple and pear. None of the materials were toxic to honey bees or reduced pollen gemination or fruit set.
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Fruit set and yield of pear cultivars as affected by reduced bee pollination period
11-16.Views:450Results of our experiments prove that pear is more or less sensitive to the reduced bee pollination period. However, the reaction (or the sensitivity) of cultivars may be different to the reduced bee pollination. Most cultivars produce much less yield under reduced bee pollination or no yield with the exclusion of bees but in the case of some cultivars total exclusion of bees does not prevent the yield formation and what is more sometimes reduced bee pollination can be resulted in somewhat higher yield than open pollination. Typical reaction, however, is a significant yield reduction with reduced bee pollination. Pear seems to be somewhat less sensitive to the partial reduction of bee pollination period than apple or quince. The first half of the flowering period seems to be more important in yield formation because usually higher yield was resulted when pear cultivars received open pollination in the first than in the second half of the blooming period. Based on our experimental results no definite relationship between parthenocarpic capacity of cultivars and the yield under reduced bee pollination can be established. So reduced bee pollination does not seem to contribute the parthenocarpic fruit formation in pear.
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Training systems of fruit trees in Hungary
123-127.Views:1102Growing sites and soil conditions of Hungary warrant profitable production of several temperate fruits at elevated levels of quality. The climate of the Carpathian basin is a mixtures of three main climatic zones the prevalence of which may change seasonally: Atlantic, continental and Mediterranean, therefore, growing sites are rather various. Temperature minima of the winter and late spring frosts are the main elements of risk. In choice of the system of cultivation, regularity of yields and intensity are to be observed equally. Regular yields are particularly aimed in stone fruit cultures.
For apple and pear plantations of high density required for intense production are promoted favourably. Accessories of intense orchards (irrigation, supporting system, rootstocks, phytotechniques, etc.) are important. In peach and plum trees are trained to funnel-shape crowns, in general, intense-types are possible in plum, only. In apricots, a Hungarian speciality, the "umbrella" type of crown is applied, almost exclusively, according to Papp. In sweet and sour cherry, the harvest technique, manual or mechanised, according to the intended utilisation, are determining the form of training.
Red and black currants as small fruits are grown mostly as bushes or hedgerows without any supporting system designed to facilitate mechanical harvest. Raspberries and blackberries are grown as hedges on trellis. Gooseberry is a special case, being a low, thorny bush difficult to be picked. Thus grafted small trees are attached to a wire-trellis which helps to solve problems of plant protection too.
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Comparative study of sour cherry samples for their anthocyanin content measured by tools of the laboratory and by portable equipments
29-31.Views:917The inner quality of fruits depends on many components. The analysis of pharmacologically important fruit species as the sour cherry consumed as a common food is an expensive task, mostly requiring an equipped laboratory. In sour cherry, one of its valuable components, the anthocyanin content deserves special attention. A portable analytical implement, which has been developed by our team measures the anthocyanin content on a scale of 1024 PharMgrades. The system is a member of the UVEX family of implements working with a microprocessor and performs the measurement quickly. The portable variant (UVEX-ML-1) does not require much training, the laboratory variant (UVEX UL-1) performs a large quantity of tests under modest laboratory conditions. Preparation of the samples is easily done, the necessary elements are available in the trade. The reagent is easily stored and dosed. The system was tested and proved to be of sufficient precision and the result showed acceptable variance according to the checks performed by a spectrophotometer in the laboratory.
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Test of the utility of apple retrotransposon insertion patterns for molecular identification of 'Jonathan' somatic mutants
7-10.Views:573Up until today, apple sport mutants proved to be indistinguishable from each other and their progenitors at the molecular level using random amplified polymorphic DNA (RAPD), amplified fragment length polymorphism (AFLP) and simple sequence repeat (SSR) marker techniques. This is not surprising, since the genomes of these somatic mutants differ only in one or a few small regions that affect economically important characteristics, such as improved fruit colour, size, or flavour. In most cases, these genome differences are probably caused by retrotransposons which are able to convert their RNA transcripts to DNA with reverse transcriptase enzyme prior to reinsertion, but unable to leave the genome and infect other cells. Retrotransposon insertions can alter the expression of other genes and/or the structure of encoded proteins. The sequence-specific amplified polymorphism (S-SAP) technique is capable of revealing the genetic distribution of retrotransposable elements over the whole genome. The present study used this approach to try to characterize and distinguish 'Jonathan' somatic mutants via fingerprinting, which is an unsolved problem.
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Growth and productivity of a young apple orchard at different spacing
23-27.Views:329Planting of new high density apple orchards showed an increasing tendency over the last ten years. Growers use in those orchards mainly dwarfing or semi dwarfing rootstocks. The spacing for those orchards is recommended based on Dutch and German experiences; however, the optimization of orchard planting density as a key factor for successful orchard management should consider the local climate conditions. An experimental orchard was planted in 2000 to investigate the effect of spacing on three dwarfing rootstocks with two apple cultivars `Jonica' and 'Gala Must'. We compared 8 planting densities (1270-3704 tree/ha) and two tree shapes (slender spindle and vertical axis). In this paper the data of the first five years' growth and bearing are presented. After 4 years, the decreasing tree densities caused reduced trunk cross sectional area. Tree density had a significant positive effect on cumulative yield per hectare. From the examined rootstocks, M.9 Burgmer 984 gave the smallest canopy for both cultivars. The trees on M.9 T.337 and on Jork 9 rootstocks have stronger growth. The light interception was measured under the canopy by AccuPAR (Decagon Devices Inc.'s).
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Relaco: a product to reduce water shoot formation around pruning wounds in pome fruit trees
30-31.Views:198Larger cuts made during the winter pruning of pome fruit orchards a number of serious problems: the wounds serve as entrances for several branch-infecting pathogens, the numerous water shoots formed around the cut edges consume assimilates, shade the developing fruits, make the canopy dense, reduce the effectiveness of the pest management and cause extra work during next seasons pruning work. These problems can be eliminated by using proper wound protectants, like Relaco, a new Romanian wound protectant to cover larger pruning cuts of apple, pear and quince trees.
The treaments by Relaco resulted in a significant decrease in both the number and total length of water shoots formed around the cut egdes.
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Integration of terrestrial laser scanning and spectral canopy scanner in horticulture applications
15-18.Views:1044One of the most difficult challenge in the everyday practice to describe the canopy growing of fruit trees in an orchard. The photosynthetic activity is the basic of the primer production of plants. The measurement of leaf area and determination of the photosynthetic activity could be occurred with some elaborated methods between experimental conditions. In this article we present such an integrated methodology, which is ideal to determine the geometric and spectral characteristic of fruit trees between field conditions.We have carried out laser scanning technology to investigate the geometric-topological characteristics and parallel the active infra-red sensor to collect spectral data about an apple orchard. The surveys were worked out in an intensive apple orchard with drip irrigation system, protected by hail net in Study and Regional Research Farm of the University of Debrecen near Pallag. This study shows the filtering and interpretation methods of created data. The produced high accuracy data can be directly used in the precision horticulture. It could serve as a guiding data to implementation a future “virtual horticulture”. Higher spatial and temporal resolution could help for a better recognition of water balance of orchards.
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Foliar application of zinc and its effect on greenhouse grown cucumber
79-82.Views:544The experiment was conducted to examine the effect of the foliar application of zinc on yield and crop quality and on fruit mineral composition of cucumber plants (Cucumis sativus cv. Mustang) which were grown in peat in containers under unheated conditions and were not showing visible signs of zinc deficiency.
In the trial the following 3 treatments were set up in 4 replications: Znl = 0.35 g/litre Zn, as foliar fertilizer; Zn2 = 0.7 g/litre Zn, as foliar fertilizer, control = no foliar fertilization. Foliar fertilization was applied 5 times with 10 day intervals. After their planting out the plants were fertigated daily with water soluble complex fertilisers. Fruits were harvested twice a week, 16 times in all, and were divided into three quality classes (class I, class II and substandard). Shoot length of the plants (plant height) was measured on 3 occasions. Zinc content of the fruits and leaves was analysed at two times.
From the results of the trials it can be concluded that the 0.35 g/litre Zn (0.35 mg/ml) foliar fertilisation had beneficial effect on cucumber both in terms of yield and quality. Under the conditions of the experiment (daily fertigation through drip irrigation) the effect of a more concentrated foliar application of zinc seemed less beneficial.
The zinc content of the fruits showed no evident increase in response to foliar fertilization, while a significant increase was seen in the leaves, particularly with the more concentrated Zn treatment. This indicates that in the case of cucumber zinc, through its assimilation in vegetative parts, has an indirect effect on fruit development.
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Experimental results of the effects of Hungarian climatic conditions to German disease-resistant industrial apple varieties
53-56.Views:206In the recent years, several disease-resistant apple varieties appeared through the modern breeding technologies. These varieties can be grown with low usage of pesticides, which mean not only environment friendly fruit growing, but the production costs are also lower. In Eastern-Hungary — it is one of the main apple growing regions — a new apple growing structure started to form by the investment of the German Wink Ltd. — several resistant apple varieties were brought from Germany. 'Resistant' refers genetic resistance that usually transferred from the genome of wild apple species. But the fruit of these apple species is not only resistant to diseases, but its quality is poorer, too. In Germany the Re-apples are grown only for the processing industry. Due to climatic circumstances in Eastern Hungary, the first experiences showed better parameters during laboratory measurement, the fruits have more beautiful view, shape and inner characters than usual industrial apples.
In our paper we discuss the results of sensory (consumer) tests, carried out in Eastern Hungary and in the Budapest-region the data analysis of systematic storing experiments (refraction, flesh firmness, weight loss, etc.) and profile analysis of fresh and stored Re-apples. (In the profile analysis the ProfiSens software [4,5] has been used.)
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Water use of young apple trees related to leaf area development
61-63.Views:389Present paper describes an investigation concerning seasonal water use and foliage area development of apple trees. Sap flow velocity was measured in the trunks of five years old apple trees cv. `Florina'/M.26 by a thermal dissipation (Granier) method from 20th of May to the end of September in 1998. The development of foliage area was estimated by a method including leaf area measurements, recording of leaves and shoot length. The foliage area reached to 70% of the maximum yearly value at beginning of June. The remaining 30% developed to the end of August. The leaf area specific water use was considerable higher in June and July, than in second part of summer. The trends of ET-FAO and water use curves differed mostly in the late season: the ET-FAO curve falls quite in September compared to August, whereas the value of water use was a similar as in August. This insensitivity of ET-FAO in this period may be a great disadvantage while using the Penman-Monteith equation in irrigation scheduling of apple.
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Sensorial appraisal of sour cherries grown by environmental technology
27-30.Views:972Sour cherry samples have been tested by a panel of potential consumers, and the judgments expressed by points given for each attribute. The correlation of individual traits with “general impression” has been explored. Primary raising of data was aimed to evaluate the effect of environmental growing technology upon sensorial impression of fruits for fresh consumption. Organoleptic tests consist of tasting and filling out of forms with numerical data as ratings them regarding taste, flavour and flesh firmness. The values of coefficients of correlation showed that a couple of other attributes as appearance, juiciness and sugar/acids perceived were irrelevant from the point of view of acceptance, i. e. the general impression.
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The hungarian raspberry today and outlooks of its development (Review)
7-11.Views:783The hungarian raspberry today and outlooks of its development (Review)
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Flower constancy of honeybees (Apis mellifera L.) to blooming pear plantations
81-85.Views:1321Studies were made on the composition of pollen loads of honeybees captured at the flowers of blooming pear trees in pear plantations. Also the foraging behaviour of honeybees was observed. Overwhelming majority of honeybees visiting the flowers of 13 pear cultivars in 1996 were pollen gatherers (95.6 per cent). Proportion of pure nectar gatherers was as low as some 3.7 per cent and no more than 0.7 per cent performed mixed behaviour. The analysis of pollen loads of bees collected at pear flowers in blooming pear plantations showed that fidelity was as high as 89-90 per cent towards pear, higher than for another fruit species in other studies. Even those plant species that are regarded to be strong competitors of blooming fruit trees in the literature (Taraxacum officinale, Stellaria media, Lamium purpureum) were scarcely represented in the loads. Accordingly, honeybees can be much more important and more effective pollinating agents of pear cultivars than generally believed.
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Preliminary evaluation of breeding perspectives of Ukrainian sweet cherry cultivars: nutraceutical properties and self-incompatibility
7-11.Views:991Some traditional sweet cherry cultivars of Ukrainian origin may represent perspective material for Hungarian cherry breeding. A total of eight cultivars analysed represent great diversity in several phenotypic traits including fruit ripening time or fruit flesh colour. Considerable differences in the anthocyanin content may result in different antioxidant capacity of fruits. In the present study, we used ferric reducing antioxidant power (FRAP) and total phenolic content (TPC) assays to characterize fruits’ nutraceutical properties. These values were compared with the respective values measured for eight commercial cultivars grown in Hungary. The average of FRAP and TPC values was higher for the Ukrainian cherries compared with commercial cultivars suggesting they might be included in functional breeding programs. Since, cherry is a self-incompatible species, the determination of S-genotype is required for both breeding and successful cultivar association in commercial orchards. Complete or partial S-genotypes were determined for 5 and 3 cultivars, respectively.
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Characterization of quince (Cydonia oblonga Mill.) cultivars using SSR markers developed for apple
7-10.Views:1202Quince (Cydonia oblongaMill.) is a minor fruit crop, which is primarily used for marmalade, jam and sauce.Very few quince cultivars are known all over the world and in many cases similar names are used for presumably different cultivars. The aim of the present study was to evaluate and characterize the genetic diversity of 36 quince cultivars and selections with SSR markers. Seven out of 8 SSR markers designed from apple sequences could successfully yield amplification also in quince cultivars. Number of alleles per locus ranged from 2 to 3 alleles. These allele numbers are quite low when compared to apple. It is supposed to be the consequence of a genetic bottleneck. In spite of the low allele number per locus, the 36 quince cultivars formed 30 different genotypes. The ratio of homozygosity was low, which might be coupled with the self-(in)compatibility phenotype of quinces. SSR markers proved unable to differentiate putatively closely related cultivars (e.g. ‘Bereczki’ and ‘Bereczki bôtermő’). In general, the level of polymorphism among the tested quince genotypes was much restricted due to the low allele number detected. However, it must be considered that the number of analysed SSR loci is not enough high to estimate the overall heterozygosity of the quince genome. Further experiments are needed and the SSR markers proved to be a reliable and useful tool for such analyses.
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Inheritance of blooming time in walnut, with regard to the property of reproductional autoregulation of species
118-122.Views:226A great number of crosses have been made with Hungarian and foreign varieties as partners to breed improved varieties. This species shows a particular trait, namely the autoregulation of fruit set, which affects considerably the productivity of commercial orchards. Thus the inheritance of the blooming time of the male and female flowers has been explored for several years in the progenies.
It has been stated that
- the feature of the partners does not turn up predictably in the progeny,
- it is most important to take into consideration the blooming time class of both, male and female flowers in planning associations of varieties for commercial orchards.
- in years of irregular spring weather the stability of the blooming time of the variety or in other words the deviation of the actual blooming time of variety from its characteristic blooming-time class is also very important.
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Blumeriella jaapii (Rehm) v. (Arx) infection of some sweet cherry cultivars in two years with different precipitation conditions
37-39.Views:295We examined Blumeriella jaapii (Rehm) v. (Arx) infection on 21 sweet cherry cultivars in the cultivar-collection of the Experimental Station Pallag of the University of Debrecen, in 2004 and 2005. Considerable differences were detected in the infection degree of the different cultivars. In 2005, due to the more humid weather, the average infection highly exceeded data of the previous year. No symptomless cultivars were detected in either year. The least sensitive ones were Linda and the cultivar 11/106. The most sensitive cultivars were Early Müncheberg, Biggareau burlat and Round Solymári. Infection degrees of Alex, Rita and Vega were different from those of other cultivars on the basis of the two experimental years.
It is important to consider both the specific weather conditions of the year and cultivar resistance at planning the plant protection system. It is recommended for each cultivar to perform a fungicide-spaying after the harvest.
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Overwintering capability and spring population size of honeybee colonies (Apis mellifera L.) in Hungary
153-156.Views:209Honeybee races and ecotypes of different genetic background have different population development in spring. Some of them can reach the necessary population size by the beginning of Robinia pseudoacacia (black locust) blooming period. There were significant differences in the spring population development between the colonies of different genetic background. The Italian races (A. m. ligustica) and their cross-breeds over-wintered poorly in Hungary, their spring population was low and they collected small amount of Robinia honey. The Austrian improved Carniolan (A. m. carnica) colonies over-wintered well, they had the largest spring population in both years. There was no significant difference between the size of the spring population of the same colonies of different genetic background in 1995 and 1996. The rate of the population development of the colonies was different in the two examined years. There was strong correlation (r = 0.8) between the spring population size and the Robinia honey yield, and between the mid-April population size and the Robinia honey yield of the colony groups of different genetic background. Spring population size also important in the effective pollination of fruit tree species that bloom earlier than the black locust trees.
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Effects of irrigation frequency and repellent plants on false codling moth (Thaumatotibia leucotreta) infestation, growth, yield, and quality of sweet pepper (Capsicum annuum L.)
34-40.Views:96Sweet pepper (Capsicum annum L.) is an important vegetable and spice crop grown worldwide for its pungency, which is derived from high concentrations of capsaicinoids. Despite the economic importance of sweet pepper as an export vegetable in Kenya, its production is currently constrained by the false codling moth (FCM) (Thaumatotibia leucotreta) and inadequate management strategies for the pest, among other factors. This study, therefore, evaluated repellent plants (Artemisia (Artemisia absinthium), Lavender (Lavandula angustifolia), and Spearmint (Mentha spicata) under different irrigation frequencies to control FCM. A split-plot factorial greenhouse experiment was conducted at Kenya Agricultural and Livestock Research Organization, Muguga (trial 1) and the Horticulture Research and Teaching Field, Egerton University, Kenya (trial 2). Irrigation frequency at three levels (irrigating once a week, irrigating twice a week and irrigating thrice a week) was the main-plot factor, while repellent plants at four levels: sweet pepper (Capsicum annum L.), artemisia (Artemisia absinthium), lavender (Lavandula angustifolia) and spearmint (Mentha spicata) constituted the sub-plot factor. Irrigating thrice or twice a week and using lavender as a repellent plant significantly reduced false codling moth infestation by 90%, increased yield by 28% and improved fruit quality in terms of fruit collar diameter and total soluble salts (TSS). The findings demonstrate that integrating lavender repellent plants with frequent irrigation can be adopted as an eco-friendly management strategy for FCM and enhancing sweet pepper yield and quality.
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Factors affecting apricot fruit antioxidant capacity and mineral element contents
95-99.Views:528Several epidemiological studies revealed that the consumption of antioxidant compounds and the risk of atherosclerosis, increased blood pressure or cancer are inversely proportional. The individual amounts and relative proportions of macro- and micro elements present in food is also of great consideration since these are involved in a wide range of physiological processes including the influence of the redox homeostasis. The antioxidant power and mineral nutrient content of fruits might be affected by several factors including genotype, ripening stage, year-effects or a wide range of environmental conditions. This study was carried out to survey the antioxidant power and mineral element content in fresh fruits of apricot and analyse some genetic and environmental factors that may have important contribution to the inner content of apricot fruits. In addition, the influencing effect of the extraction procedure used for antioxidant analyses was also tested. Our analyses indicate that a considerable fraction of antioxidant capacity is attributable to the hydrophilic antioxidants. The genetic background has crucial importance in determining apricot fruit antioxidant capacity and mineral nutrient content; however, the growing season and the ripening time of fruits may have also important effects.
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Situation of peach resistance to diseases in Romania
65-69.Views:342The resistance of peach cultivars to the most important diseases caused by fungi, bacteria, viruses and mycoplasms was studied in Romania over two working stages in the period between 1985-2005. The major diseases examined were: Cytospora cincta Sacc., Taphrina deformans (Berk) Tul., Moruluua laxa (Aderh. & Ruhl) Honey, Sphaerotheca pannosa var. persicae Woron., Stigmina carpophila (Lev) M.B.Ellis, Pseudomonas syringae pv. syringae, mycoplasm and plum pox potyvirus. Based on the obtained results, the studied cultivars were classified into resistance groups for the different diseases (very resistant, resistant, medium resistant, sensitive and very sensitive).Based on the results of our study, the following gene sources were chosen following the evaluation of the various genetic material in the peach germoplasm fund, in the climatic conditions of Romania: Cytospora cincta: Cullinan, Cardinal, Hamlet, NJF 3, Onakita Gold, Triumf, "Superba de Toamna", Anderson, Weinberger; Stigmina carpophila: Armgold, ARK 109, Stark Early Blaze, Cardinal, Congres; Taphrina deformans: Madeleine Pouyet, Cumberland, Harbelle, Indian Blood, Sulivan, Victoria, Zafrani, Pekin, Naradnji Ranhii; Spaherotheca pannosa var. persicae: Triumf; Congres; Victoria; Armking; Morton; Regina; Nectared 7; ARK 125; ARK 134; Regina.
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Comparative study of plum cultivars belonging to different taxons during 1980-1996
13-19.Views:307The program of the Research Institute of Cegléd, comprises the acclimation and selection of 7 Prunus-species and 39 cultivars grafted on Myrobalan C. 679 (P. cerasifera) seedling stocks. In addition two varieties have been investigated on three different rootstocks each: Myrobalan C. 174 (P. cerasifera), Bitter almond C. 449 (P. arnygdalus var. amara) and Sweet almond C. 471 (P. amygdalus var. dulcis). Fruits of commercial quality are produced maroly on some cultivars of P. salicina-, P. italica- and P. cerasifera character. Authors explain the three possible causes of low yields experienced in non-European plums:
1/ unfavorable environmental conditions
2/ lack of irrigation and
3/ superficial information concerning the variety, rootstock and adequate traditions as well as growing practices.
A rather tight correlation has been stated between blooming dates and the main ripening period. However, the early blooming time alone cannot be considered as the cause of low productivity. The decay of plum trees is attributed to special ecological requirements and phytosanitary problems of the foreign plum cultivars. The analysis of regression revealed stochastic relations involving several other characters too, which facilitate the planning of cross-combinations in the breeding program.
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Incidence of brown rot blossom blight caused by Monilinia laxa in organic sour cherry production in Hungary
133-135.Views:617Brown rot blossom blight incidence was evaluated on two sour cherry cultivars (Kántorjánosi, Újfehértói fürtös) in an organic sour cherry orchard in Hungary in 2003 and 2004. Trees were grown according to the organic fruit production guidelines. Blossom blight incidence was similar in both cultivars in both years. In 2003, blossom blight incidences were low even in the untreated plots (less than 15 %). In 2004, brown rot blossom blight incidence reached almost 30 % in the untreated plots. Blossom blight incidence was significantly lower in the conventionally treated plots in both cultivars.
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Anatomical relations of root formation in strawberry
71-75.Views:509Anatomical relations of root formation are traced throughout the life cycle of the strawberry plant from the germinating seed up to the runners of the adult plant. Histological picture of the root changes a lot during the development of the plant. First the radicle of the germ grows to a main root, which makes branches into side roots and later adventitious roots are formed on the growing rootstock or rhizome. The anatomy of the different types of roots is also conspicuously different. First tiny branches appear relatively early after germination on the seedling's radicle, but soon the hypocotyl of the seedling thickens and develops side roots, which are already somewhat stronger. During this interval, the first true leaves are formed. The 4th or 5th of them being already tripartite, and the initiation of new roots extends into the epicotylar region of the shoot. The second years growth starts with the development of reproductive structures, inflorescences and runners starting from the axils of the new leaves. Near the tips of the runners below the small bunch of leaves, new root primordia are initiated. The tiny radicle of the germ develops a cortical region of 5-6 cell layers. Cells of the central cylinder are even smaller than the cortical parenchyma and include 3-4 xylem and 3-4 phloem elements as representatives of the conductive tissue. Roots originating from the shoot region are much more developed; their cortical zone contains 17-20 cell layers, whereas the central cylinder is about half as large. In the next year, new roots are formed at the base of the older leaves. These roots differ hardly from those of the last season in size and volume, however, they are recognised by colour and their position on the rhizome. The roots of the last year are dark, greyish-black, and grow on the lower third length of the rhizome, on the contrary, the new ones, on the upper region, are light brown. Roots starting from the shoot or rhizome are, independently from their age or sequence, mainly rather similar in size and diameter, thus being members of a homogenous root (homorhizous) system, i.e. without a main root. Plants developed and attained the reproductive phase develop in the axils of the leaves runners being plagiotropic, i.e. growing horizontally on the surface of the soil. The runners elongate intensely, become 150-200 mm, where some long internodes bear a bunch of small leaves and root primordia on short internodes and a growing tip. Runners do not stop growing, generally, further sections of 15-25 cm length are developed according to the same pattern, with small leaves on the tip. The growing tip of the runners is obliquely oriented, and small, conical root primordia are ready to start growing as soon as they touch the soil. The roots penetrate the soil, quickly, and pull, by contraction, the axis of the runner downwards, vertically, developing a new rhizome. The short internodes elongate a little and start developing adventitious roots. At the end of the growing season, the plantlets arisen on the rooted nods of runners are already similar to the original plants with homogenous root system. On the side of the adventitious roots, new branches (side-roots) are formed. The root-branches are thinner but their capillary zone is more developed being more active in uptake of water and nutrients. The usual thickening ensues later.
