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Contributions to the flora of kurgans in the Middle Tisza region
94–105Views:552Kurgans are ancient burial mounds built by nomadic steppic cultures. Embedded in the heavily transformed landscapes of the Carpathian Basin they often serve as last refuges for rare and endangered plant species. In our paper we publish floristic data collected on 82 kurgans located in the area of the Hortobágy National Park Directorate. Our dataset covers 64 CEU quarter quadrates, and the territory of 45 settlements. We provide data on 39 taxa that are either protected (e.g. Anchusa barrelieri, Centaurea solstitialis, Ranunculus illyricus and Phlomis tuberosa) or regionally rare (e.g. Aegilops cylindrica, Astragalus austriacus, Glaucium corniculatum and Trifolium diffusum). Our records demonstrate that kurgans have a vital role in maintaining the populations of rare and endangered grassland species even in transformed landscapes.
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Botrychium lunaria in the Mecsek Mts
91-92Views:260A single individual of Botrychium lunaria (L.) Sw. was found in a shady dolomite rocky grassland above the city of Pécs in the Mecsek Mts., on 1 June 2020. The habitat of this fern, which is new to the flora of the Mecsek Mts., is presented by a phytosociological relevé.
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Distribution of Campylopus introflexus (Hedw.) Brid. in Hungary
212-219Views:117Based on field work data collected in 2013–2014, the authors generated a map of the current distribution of Campylopus introflexus for Hungary. Besides the coordinates and geographic name of the localities, an estimated size of the population, the type of habitat and substrate, the co-occurring moss species and affected vegetation types were also recorded. Altogether, 18 stands were found in 8 geographical regions of Hungary, in some cases Campylopus introflexus was found to be a new species record for the moss flora of the respective region (e.g. Dunántúli-dombság, Nyugatmagyarországi peremvidék). In addition to the pine plantations from where it was detected formerly, the mixed deciduous-coniferous forests, the acidophilous oak woodlands and man-made habitats were found to be also proper habitats for this moss taxon. Earlier data mainly originated from pine deadwood, the recently discovered stands were found on acidic soil surfaces. The most important cooccurring moss species were the followings: Hypnum cupressiforme, Polytrichum piliferum, P. formosum, Pohlia nutans and Ceratodon purpureus. Campylopus introflexus had prominently high coverage in the acidophilous oak forests of the Mecsek Mts, where it occurs on acidic sandstone covered with a thin debris of the bedrock. This neophyte moss is widely distributed in Hungary, and has stable populations. Further expansion is expected in the subatlantic and submountain regions of the country, and in regions characterised by acidic soils and pine plantations.
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Vaccinium microcarpum (Turcz. ex Rupr.) Schmalh. in Hungary
71–76Views:117The plant that was reported as Vaccinium oxycoccos L. from the Nyírjes-tó (Sirok village, NE Hungary) in 1994 was subsequently (1999) identified as Vaccinium microcarpum (Turcz. ex Rupr.) Schmalh. by the first author of this article. To substantiate this identification morphometric data were collected and analysed from Vaccinium specimens occurring in the Sphagnum dominated mires of Nyírjes-tó (Sirok vilage) and Nyíres-tó as well as Báb-tava (Bereg-plain) in September 2016. Field photographs taken previously at the localities of Bereg-plain were also used for identification. The leaves of the Sirok plants are smaller, ovate-triangular, i.e. widest near base. The ovate-elliptic bracteoles are inserted on the basal third of the glabrous pedicels. The flowers are solitary, and have dark pink corollas and hairy filaments on the whole abaxial surface. The fruit is pyriform. In contrast to the Sirok plants, those of the Bereg-mires have clearly larger and ovate-elliptic leaves. The filiform bracteoles are inserted on the middle third of the minutely hairy pedicels. The whitish flowers are grouped in clusters of (2)–3–4–(5), i.e. they form an inflorescence. The filaments are hairy only on the margins of the abaxial surface. The fruit is globular. The statistical analysis confirmed that the Sirok plants belong to V. microcarpum, whereas the Bereg-plain plants represent V. oxycoccos.
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New occurrences of Ranunculus strigulosus in East Hungary
3–8Views:360The exact distribution of Ranunculus strigulosus Schur is poorly known in Hungary. Current occurrence data of the species mainly from Békés county and only one archive data from the Nyírség region, from the 1920s were known. This study presents eleven newly discovered occurrences in East Hungary. Populations were found in wet meadows, forest glades and poplar plantations. Nine of these populations are located at the southeastern edge of the Nyírség region and most of them are quite small. Three larger population were found in Nyírábrány, Álmosd and the Tócó Valley between Debrecen and Hajdúböszörmény. One of the newly discovered populations provides a new contribution to the flora of Bátorliget Marsh and Fen-wood NCA. Some morphological characters (hairs on the stem and the leaves, shape of the leaves developed before and after the mowing) are also discussed.
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Data on the distribution of some protected plant species and on their presence in secondary habitats
183-199Views:1828In this paper, I publish my floristic data on some protected and strictly protected plant species collected in Hungary between 2016 and 2021. The data published in this paper cover 31 micro regions of Hungary, but most of them come from the Great Hungarian Plain, especially from Pest and Bács-Kiskun counties. I describe the occurrences of 41 protected and three strictly protected vascular plant species (two ferns and 42 flowering plants), some of which represent new data for the particular micro region or even larger geographical areas (e.g. Botrychium matricariifolium, Thlaspi alliaceum and Nasturtium officinale). For some species, the published data either come from floristically understudied areas (e.g. Ranunculus illyricus, Vinca herbacea and Cardamine amara) or are considered rare in the certain region and/or in the whole country (e.g. Apium repens, Cnidium dubium and Vicia narbonensis). For other species, the newly reported occurrences are presented as complement to the already known distribution data (e.g. Cephalanthera damasonium, C. rubra, Linaria biebersteinii, Sonchus palustris and Agrostemma githago). In support of the nature conservation actions, I provide data from the flora of some floristically understudied micro regions (e.g. Dorozsma-Majsai Sand Ridge, Bácskai Loess Plain) and try to highlight the role of canals in Hungary in species and biodiversity conservation, by the example of the Danube-Tisza canal. In this manuscript, I publish data on twelve protected plant species from the Danube-Tisza canal and its canal bank (eg. Peucedanum palustre, Carex paniculata, Clematis integrifolia).
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Contributions to the flora of the Danube River in Komárom-Esztergom and Fejér counties (Hungary)
157–168Views:289The present paper focuses on the aquatic and Nanocyperion vegetation of the Danube floodplain. Most of the data show the results of the surveys in 2019 at low water levels. The Nanocyperion and aquatic vegetation of the Komárom-Esztergom County section is relatively well researched compared to the Fejér County section which was previously poorly known. In the last decade, two rare species – Hippuris vulgaris and Hydrocharis morsus-ranae – have disappeared between Gönyű and Neszmély (Komárom-Esztergom County). In contrast, several new invasive plants have appeared: Salvinia molesta, Myriophyllum aquaticum, Limnobium laevigatum, and Crassula helmsii (first record from Hungary). Vallisneria spiralis and Elodea canadensis were detected again after several decades. Elodea nuttallii and Azolla sp. have become more widespread and abundant than previously. New and uncommon Nanocyperion species were the native and protected Lindernia procumbens and the invasive Lindernia dubia. New invasive plants in the studied Danube sections were: Cyperus esculentus, Euphorbia maculata, and Eleusine indica. The native Veronica catenata and Schoenoplectus triqueter have become more widespread than before. Polygonum graminifolium was an interesting find of open pebble surfaces in several parts of the examined area, which has no recent records in Hungary from before 2015 (published here).
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The biases of floristic surveys I. Naive planters: “cuckoo” of the Medves (N Hungary)
207-217Views:145The aim of the planned thematic studies is to point out the main biases and/or errors of floristic surveys in Hungary. In this first article of the series we focus on those examples when native species were planted out of their original habitats and/or range by a non-professional naturalist. The sources of information are those letters that were posted by Ferenc Varga to the first author of this article in 2005. The circumstances of the transplantation of 10 phytogeographycally significant species of the investigated area and/or the Pannonicum (Adonis vernalis, Arum orientale, Asarum europaeum, Genista germanica, Phragmites australis, Pulsatilla grandis, P. nigricans, Salvia aethiopis, Viola alba, Xeranthemum annuum) are detailed in the text and summarised in a table. The correspondence between the first author and Ferenc Varga provides insight how the ambition of a non-professional naturalist can influence the flora of an area. Most of the species were removed from South and transplanted to North, almost 10 km away from their original stands. With the exception of Adonis vernalis and Genista germanica at least one transplantation of the species was successful. In the case of Salvia aethiopis, the naturalized populations came 1.25 km close to Slovakia, where this species is a very rare and endangered (EN) taxon.
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Contributions to the algal flora of a recently-formed peat moss bog near Kőszeg (W Hungary)
113–130Views:388The Alsó-erdő peat bog near Kőszeg has undergone a significant transformation since its first mention in the literature. In addition to the original single patch, a smaller one appeared a few decades later. The process did not stop and a third unit, discussed here, was formed. The succession of its macrovegetation towards the typical peat moss association started slowly but surely, due to balanced water regime and other optimal environmental conditions. The process is more advanced for the algal vegetation, which shows the characteristics of peat bogs already. The high species diversity found makes the habitat highly valuable itself. Although a special, unique algal species has not been found in previous studies, 27 Desmidiales taxa listed as endangered occur sometimes with high density and make the habitat even more valuable. Based on our results, the area deserves special attention both from the habitat conservation and research point of view. At the same time, its further monitoring can provide crucial information for exploring and understanding the succession of similar habitats, and thereby for their more efficient protection.
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Sorbus udvardyana Somlyay & Sennikov in the eastern part of the Balaton Uplands
55-58Views:293Related to distribution of S. aria s.l. × S. torminalis hybrid taxa in the eastern part of the Balaton Uplands just data of „S. balatonica” is in the literature based on collection of Ádám Boros (Kárpáti 1960). In the summer of 2014 the author found and mapped a few (sub)populations of S. aria s.l. × S. torminalis at Lovas (Királykúti-völgy, Atya-hegy, CEU: 8973.4) and Felsőörs (Malom-völgy, Kopasz-tető, CEU: 8973.4) villages. Based on leaf-morphological traits the specimens were identified as S. udvardyana.
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Botanical comparison of man-made landforms in the Nagykunság and Nagy-Sárrét regions (E Hungary)
65-76Views:213Loess grasslands are among the most important vegetation types of the Great Hungarian Plain. This paper compares the composition of vegetation situated on three types of man-made landforms (burial mounds, Devil’s dykes and river dikes) which were built in different historical times. I studied the similarities and differences in their vegetation, focusing on the rare species and plant associations. I studied six burial mounds, two Devil’s dykes and four river dikes. All of the studied landforms were covered by loess steppes, loess cliffs or Artemisia salt steppes. Devil’s dykes had the most valuable vegetation and they preserve a very diverse flora in the landscape.
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Interesting moss and liverwort records from the Western Mecsek Mts (South Transdanubia, Hungary)
29-38Views:107During an extensive field work in the acidophilous forests and fens of the Western Mecsek Mts. current localities of several species, which haven’t been recognised here for 50 years, were revealed (e.g. Bazzania trilobata, Diphyscium foliosum, Scapania nemorea). The authors also found some new stands of regionally and/or locally rare taxa (e.g. Buxbaumia aphylla, Dicranum tauricum, Jamesoniella autumnalis, Lepidozia reptans, Sphagnum squarrosum) as well as species new to the moss flora of the Mecsek Mts. (e.g. Aulacomnium androgynum, A. palustre, Calliergon cordifolium, Hedwigia stellata). Both investigated habitat types were found to be infected by invasive moss (Campylopus introflexus) and liverwort (Riccia rhenana) species. It is noteworthy that most of the species were found in winter or at the beginning of spring, when most vascular plants are inactive and their coverage is negligible. In this period moss and liverwort taxa are active, and are therefore easily noticeable. The authors raise the possibility that the expansion of Dicranum tauricum in Hungary is a consequence of former coniferous (especially Pinus spp.) plantations, since the old, dead trunks of pine species can serve as stepping stones to colonization for this taxon.
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Data to the flora and vegetation of Hungary III.
197-206Views:177We report the occurrence data of 44 vascular plant taxa collected at various localities in Hungary (mainly the Danube–Tisza Interfluve and South Transdanubia) during floristic, vegetation and ecological field surveys between 2016 and 2018. Of the species in the present article, 27 are protected or strictly protected. In the paper we focused on plants that are rare regionally (e.g. Botrychium lunaria, Polystichum aculeatum) or in the whole country (e.g. Ophrys oestrifera, Ranunculus psilostachys), as well as on plants that occurred in unusual habitats (e.g. Corydalis solida). We also included new localities of weedy species, some of which (e.g. Opuntia humifusa, Sporobolus cryptandrus) may present nature conservation threats.
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Data to the flora of Hungary I.
254-259Views:80The paper gives new floristic data from several regions of Hungary. Some of them are recent evidences of old data (e.g. Herniaria incana in Pomáz), others are new occurrences of recently expanding alien species (e.g. Eleusine indica, Geranium purpureum, Typha laxmannii), casual species (e.g. Lobularia maritima, Nepeta racemosa) or legally protected taxa (e.g. Agrostemma githago, Ornithogalum brevistylum, Ranunculus illyricus, Sonchus palustris).
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Results of floristic and phytogeographical research in the Velence Hills (Hungary, Transdanubian Mts)
117–152Views:436A recent floristic mapping of the Velence Hills has clarified the local distribution of many plant species. This study presents records of 219 phytogeographically significant or sporadic/rare taxa (216 species/subspecies and 3 natural hybrids). The local distribution of some species representing typical regional distribution patterns (Alyssum turkestanicum, Artemisia austriaca, Bupleurum pachnospermum, Doronicum hungaricum, Hypericum elegans, Lathyrus sphaericus, Minuartia viscosa, Ornithogalum comosum, Scabiosa canescens, Sedum caespitosum, Sternbergia colchiciflora, Vinca herbacea) are shown on maps. More than fifty taxa previously unknown from the area were recorded during the floristic mapping, including phytogeographically significant ones, such as Bassia laniflora, Brassica elongata, Bupleurum praealtum, Centunculus minimus, Hypericum elegans, Orlaya grandiflora, Phleum bertolonii, Silene dichotoma, Stipa tirsa, Tordylium maximum, Trifolium diffusum, T. ochroleucon and Valeriana dioica. In terms of phytogeography, the Velence Hills can primarily be characterised by the co-existence of continental and sub-Mediterranean elements as well as species of acidophil and strongly basic habitats.
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A new, remarkable population of Epipactis placentina found near Parádsasvár village (Mátra Mts, NE Hungary)
247–248Views:238The present paper surveyes the Hungarian records of Epipactis placentina Bongiorni & Grünanger, which is the rarest Epipactis species in Hungary. Currently, E. placentina is known only at Parádsasvár village in the country. This population (ca. 40 generative individuals) was found in 2019, in an old beech forest, in northern exposition.
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New montane species in the Hungarian bryophyte flora
16-26Views:681During a bryofloristical investigation in the vicinity of Ómassa (Miskolc) in the Bükk Mountains (NE Hungary), three montane-boreal bryophyte species new to the Hungarian bryoflora were collected. Encalypta spathulata Müll.Hal. was found on the ledges of limestone cliffs of Vörös-kő Hill as well as on the rocky debris among roots of a fallen tree lying under the cliffs. Hydrogonium croceum (Brid.) Jan Kučera and Orthothecium rufescens (Dicks. ex Brid.) Schimp. were found in the crevices of dolomite cliffs on the north-facing steep slope of Jávor Hill. While the latter two species are widespread in European Mountains, E. spathulata is considered to be a rare species, and is assessed as ‘Vulnerable’ in the IUCN red list.
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Campylopus flexuosus (Hedw.) Brid.: a moss new to the Hungarian bryophyte flora
22-28Views:109Two localities of Campylopus flexuosus new to Hungary were discovered by the authors near Uzsa in the Bakony Mts during mapping of the Hungarian bryoflora. In this paper the circumstances of the occurrences are described and illustrated by maps and photographs; a short description of the species is provided together with photographs of diagnostic characters.
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The herbarium of the Botanical Garden of Eötvös Loránd University (BPU)
55–59Views:259The paper introduces the herbarium of Eötvös Loránd University (BPU), currently stored at the Botanical Garden of the University, according to its state in the year 2013. The BPU herbarium consists of ca. 16 000 specimens the relevant data of which have been organised into an OpenOffice Calc spreadsheet database. The specimens were collected in 16 European countries, mainly in the current territory of Hungary (80%), Romania (12%) and Slovakia (5%). Other countries are represented with very few specimens. The vast majority (ca. 70%) of the Hungarian specimens were collected in Pest, Veszprém, Borsod-Abaúj-Zemplén, Hajdú-Bihar and Szabolcs-Szatmár-Bereg counties. Although the gatherings cover a century and a half period of time, the most rapid growth of the collection took place from the 1930s to the 1960s. The most prolific collectors were Rezső Soó and his followers: Lajos Felföldy, Tibor Simon and Szaniszló Priszter. Felföldy and Simon enriched the collection through more than 60 years. Data on the native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).
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Survey on the bryophyte flora of the kurgans along the river Hernád (Hungary)
153–161Views:428This paper presents the results of the first bryofloristic survey in kurgans from North East Hungary. Altogether 41 taxa (1 liverwort and 40 mosses) were recorded from nine burial mounds. The most frequent mosses were Brachythecium rutabulum, Tortula acaulon var. acaulon and Barbula unguiculata from the localities. I found several infrequent bryophyte species on the surveyed Hungarian kurgans, such as Acaulon muticum, Acaulon triquetrum, Pterygoneurum subsessile, Riccia ciliata, Tortula caucasica, Trichostomum brachydontium and Trichostomum crispulum. There was no correlation between the number of bryophytes and the area of the kurgans. The proportion of the annual shuttle life strategy, i.e. species with a very short life span, is relatively high compared to other recent studies from Hungary (e.g. Buda arboretum and Balaton village), thereby these habitats may be potential refuges for ephemeral bryophytes.
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The flora of Bolondvár (Central Hungary, Colocense)
133-141Views:108A 0.26 km2 sized Natura 2000 area called Bolondvár, located in Central Hungary, near the village Mezőfalva was studied. It is used regularly as a meadow or sometimes as a pasture. Floristic data were collected between 2010 and 2013. The total number of taxa recorded was 362. The species diversity was high compared to other places of Hungary. There were 11 protected (e.g. Ajuga laxmannii, Astragalus asper, Cirsium boujartii, Inula germanica) and numerous locally rare species (e.g. Hieracium densiflorum, Lavatera thuringiaca, Orobanche lutea, Veronica austriaca).
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Epipactis exilis in the Börzsöny Mts (C Hungary)
102-103Views:228Seven specimens of Epipactis exilis has been found in the Börzsöny Mts, in a beech forest along the creek Szén-patak on the 21st August 2020. This strictly protected species is very rare in Hungary, the present observation is its fifth occurrence in the country. It is new to the Börzsöny Mts flora.
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Contributions to the flora of Baranya and Tolna counties II.
243-253Views:74The present study reports the occurrence data of some rare and legally protected vascular plant species, collected at Tolna county and the northern part of Baranya county during field work between 2012–2013. Among the reported floristic data, the occurrence of Ranunculus lingua, Potentilla rupestris and Malva alcea are new for Tolna county. Further rare species such as Allium angulosum, Althaea hirsuta, Erodium ciconium, Galium rubioides, Glycyrrhiza echinata, Lathyrus palustris, Lathyrus sphaericus, Myagrum perfoliatum, Salvinia natans, Urtica kioviensis and Vicia lutea were also registered on the studied area.
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Asparagus verticillatus L. in Hungary
38-43Views:183Climbing asparagus (Asparagus verticillatus L.) a new adventive species to the Hungarian flora was recently discovered in the city of Pécs (South Hungary) in 2011–2012. Taxonomy, morphological description and current localities of the species are presented in this paper. A. verticillatus is a mediterranean-submediterranean species native in Balkan Peninsula, Eastern Europe, Western- and Middle Asia. The species is a popular ornamental plant in Hungary, but no data was available about its escape or naturalisation in the country so far. Herbarium specimens from Hungary were collected so far mostly from botanical and private gardens; only one specimen was collected in a dry grassland along a roadside in the city, respectively. The newly discovered small populations are located mostly in secondary dry grasslands, at the edges of vineyards and private gardens on the southern slopes of the Mecsek Mts and only one specimen was found far from the residential area in natural vegetation. Because climbing asparagus is a frequently planted ornamental plant in several locations of Pécs, further escape, naturalisation and persistence of the species is possible in the future.
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Floristic data from Lengyel–Hőgyész Natura 2000 site (SW Hungary)
31–38Views:168This paper presents occurrence data of 169 taxa from the Lengyel–Hőgyész Natura 2000 site (HUDD20026) which is on the southern part of Tolna Hills, SW Hungary. Data were collected during three years (2015, 2016, 2017), on different sites annually. The presented data contribute to the distribution dataset of Atlas florae Hungariae. Regionally rare or sporadic species (e.g. Asplenium scolopendrium, Asplenium trichomanes, Polystichum setiferum, Doronicum hungaricum, Paris quadrifolia, Scilla vindobonensis) are enumerated. Presence of Epipactis leptochila subsp. neglecta and Platanthera chlorantha were not registered from this site before. Subspecies of Carex divulsa are also studied on this site – distribution of these infraspecific taxa have not been mapped recently in Hungary.