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Data to the flora of Heves–Borsod Plain I. Distribution of forest, forest steppe and steppe elements
16-65Views:361This paper reports new floristic data of 159 taxa (158 species and one hybrid) from the Heves–Borsod Plain (5 micro-regions between the Mátra and Bükk Mountains and the Tisza River valley). The data were collected between 1999 and 2018 and supplemented with collected specimen (altogether 93 herbarium sheets). This study is the first part of a thematic series that analyzes the distribution patterns of forest, forest steppe and dry grassland ‘steppe’ species, beyond reporting floristic data. The data were derived from 51 flora-mapping quadrats (CEU), based on 5.395 field collected data records. Some indicator species of the three species groups have been assessed in detail (with the addition of distribution maps of 25 species). I have also tested two East-West direction ‘lines’ (the 100-meter isoline and the Csörsz Ditch) for the possible existence of North-South chorological gradients in the area. True forest species (mostly Querco-Fagetea elements) show a dispersed pattern, their representatives can be linked to forest blocks appearing in the landscape. Streams from the adjacented hilly areas (even if they are strongly modified) are very important in the dispersal of the forest species. Their significance and role are decreasing towards the South. For some steppe and forest steppe species (e.g. Brachypodium pinnatum, Campanula bononiensis, Clematis recta, Elymus hispidus, Lychnis viscaria, Ranunculus illyricus, Sanguisorba minor, Stipa spp., Teucrium chamaedrys, Trifolium alpestre, Vinca herbacea) the examined lines indicate a regional area boundary, while for other species (e.g. Phlomis tuberosa, Thalictrum minus) gradient-like distribution differences were not observed. In the distribution of many species an additional southern gradient running along the northern boundary of the former Heves Floodplain was detected. Southwards to this line, the representatives of the selected species are already very sporadic, occurring only in synantropic habitats (e.g. Vincetoxicum hirundinaria).
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Flowering phenology and distribution pattern of Lizard Orchids (Himantoglossum)
157-167Views:275Hybridization is very rare between the species of Himatoglossum genus, despite the fact that it is common in other orchid genera. The reasons for this can be the lack of overlapping distribution areas or the different flowering period of the species. In the paper distribution and flowering data of Himantoglossum species are presented from herbarium sheets or references (sources paper-based or internet). We collected 1001 precisely dated flowering records of 12 species. In case of 6 species (H. adriaticum, H. caprinum, H. comperianum, H. hircinum, H. jankae, H. robertianum) we had sufficient records to calculate the average flowering day and blooming length. According to the collected data the distribution of the species are overlapping except for H. formosum, H. galilaeum and H. metlesicsianum. Studying 55 species pairs, insignificant differences in blooming time were only at 21 species pairs, among them only 5 (H. comperianum - H. caprinum, H. montis-tauri - H. comperianum, H. hircinum - H. adriaticum, H. montis-tauri - H. caprinum és H. jankae - H. calcaratum) have overlapping distribution area. Data on hybridization could be found only at three species pairs among them. From the results we can conclude that in the case of Himantoglossum genus the rarity of hybridization is mostly due to the phenological isolation.
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Distribution and infection rate of (Viscum album subsp. austriacum) in Hungary
89-102Views:294Based on the data collected between 2015 and 2023, we present a detailed account of the occurrences and current distribution of Viscum album subsp. austriacum in Hungary. We present its frequency and abundance, and also examine the factors influencing its distribution and prevalence. The taxon is generally widespread in the western half of Western Transdanubia, where it is a characteristic (and partially native) element of the mixed pine forests of the Vendvidék, Felső-Őrség, Kőszeg Hills, and the Pinka Plain. Its occurrence has been confirmed in almost all (older) Scots pine and black pine stands of the Sopron Hills, where it has become a mass pest in several locations. Based on our studies, the current distribution of the taxon in Southwestern Transdanubia follows the border of the native distribution of Scots pine, but in the Kemeneshát and the Sopron–Vas Plain, the regional boundary extends 10–25 km beyond it. The frequency of pine mistletoe within the subcompartments was highest in mixed coniferous stands. We also recorded a mass infection that played a significant role in the degradation of the stands. The notable increase in the number of occurrences can be explained by the significant under-mapping of the taxon to date, as well as its documented spread throughout Europe. It is also assumed to be linked to the increasingly unfavorable health status of coniferous forests in Hungary. This process may be facilitated by the fact that the resistance of Scots pine forests in both hilly and lowland regions has significantly declined due to the increasing length and frequency of drought periods. Compared to previous years, forest stands in weakened condition are more susceptible to infections by pathogenic organisms, such as mistletoe.
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Results of floristic and phytogeographical research in the Velence Hills (Hungary, Transdanubian Mts)
117–152Views:657A recent floristic mapping of the Velence Hills has clarified the local distribution of many plant species. This study presents records of 219 phytogeographically significant or sporadic/rare taxa (216 species/subspecies and 3 natural hybrids). The local distribution of some species representing typical regional distribution patterns (Alyssum turkestanicum, Artemisia austriaca, Bupleurum pachnospermum, Doronicum hungaricum, Hypericum elegans, Lathyrus sphaericus, Minuartia viscosa, Ornithogalum comosum, Scabiosa canescens, Sedum caespitosum, Sternbergia colchiciflora, Vinca herbacea) are shown on maps. More than fifty taxa previously unknown from the area were recorded during the floristic mapping, including phytogeographically significant ones, such as Bassia laniflora, Brassica elongata, Bupleurum praealtum, Centunculus minimus, Hypericum elegans, Orlaya grandiflora, Phleum bertolonii, Silene dichotoma, Stipa tirsa, Tordylium maximum, Trifolium diffusum, T. ochroleucon and Valeriana dioica. In terms of phytogeography, the Velence Hills can primarily be characterised by the co-existence of continental and sub-Mediterranean elements as well as species of acidophil and strongly basic habitats.
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Contributions to the Atlas Florae Hungariae XIII.
85-88Views:58403The present paper is the 13th in a series of papers contributing new floristic data to complement the distribution maps of Atlas Florae Hungariae. We present altogether 1307 new occurrence data to advance our knowledge of the distribution of vascular plants in Hungary. New data are presented for 634 vascular plant species from 256 flora mapping quarter quadrates (CEU). Due to the unevenness of data collection, the data are highly dispersed over the area of Hungary. Most of the presented occurrence data are from the Great Hungarian Plain, the North Hungarian Mountains, and the Little Hungarian Plain regions. The enumeration includes rare native species (e.g., Androsace maxima, Erysimum crepidifolium, Orobanche gracilis, Peucedanum palustre, Pulsatilla zimmermannii, Scopolia carniolica, Vicia biennis), rare or data deficient adventive species (e.g., Euphorbia prostrata, Euphorbia serpens, Lindernia dubia), subspontaneous occurrences of native species (Asplenium scolopendrium, Daphne laureola, Chenopodium opulifolium), species spreading along linear infrastructure (Eleusine indica, Plantago coronopus, Sorghum halepense, Spergularia salina, Tragus racemosus), and also common native species with data deficient distribution maps (Agrostis stolonifera, Bromus benekenii, Bromus japonicus, Carex caryophyllea, Carex divulsa subsp. divulsa, Equisetum ramosissimum, Vulpia myuros).
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Contributions to the Atlas Florae Hungariae V.
383–403Views:452The current paper is the 5th one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Several distribution data of 360 plant taxa from 66 flora mapping units are published in this study. Most of them are from the South Transdanubia. As a result of systematic surveys, more than 30 new species were found in 6 flora mapping quadrates. Most of the species are frequent, treated usually as weeds, but some of them are rare (e.g. Anchusa azurea, Centunculus minimus, Euphorbia stricta, Hypericum humifusum, Sagina apetala), and in some cases they are legally protected species in Hungary (e.g. Asplenium adiantum-nigrum, Dryopteris affinis, Festuca dalmatica, Gymnocarpium dryopteris, Urtica kioviensis).
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Data to the distribution and nature conservation of Sorbus bakonyensis
229-233Views:1519Sorbus bakonyensis (Jáv.) Jáv. (syn. S. majeri Barabits) is an apomictic species with a very narrow distribution range in the Bakony Mts (Hungary). Its previous occurrence data were published only from Kopasz Mt. (~Kápolna Hill, Csordás valley) above the village Márkó. It was possible to slightly expand the known area of the species to the East and to the West (Kis-Bükk Mt) of Kopasz Mt. with a detailed mapping. Due to the number of known individuals (six mature trees and a variable number of seedlings) and its small distribution range (~2 km2) the species is critically endangered. Regarding its small population, the high number of big game (wild boar, red deer) and the more and more frequent drought periods are significant risk factors. In situ preservation of the species is still a realistic goal, but ex situ propagation in tree nurseries and reintroduction to the suitable habitats of the mapped area (clearings, forest edges) are also justified.
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Contributions to the Atlas Florae Hungariae III.
122-146Views:1513The current paper is the third one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 558 vascular plant taxa from 186 flora mapping quadrates (CEU) are presented. New records are indicated from different regions of Hungary; however, most occurrences are located in the North Hungarian Mts. Occurrence data of rare native taxa (e.g. Marsilea quadrifolia, Salicornia prostrata, Stellaria palustris, Potentilla patula, Althaea cannabina, Lythrum tribracteatum, Ajuga laxmannii, Plantago schwarzenbergiana, Alisma gramineum, Gagea bohemica, G. szovitsii), rare or data-deficient alien taxa (e.g. Chorispora tenella, Thladiantha dubia, Senecio vernalis) as well as frequent but more or less underrepresented taxa (e.g. Scleranthus annuus, Vicia hirsuta, V. lathyroides) are also enumerated. In case of the most common species we provided the CEU codes only.
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Contributions to the Atlas Florae Hungariae X.
101–106Views:682This paper is the 10th in the series aiming to contribute with new distribution data to the maps published recently in Atlas Florae Hungariae. Distribution data of 686 plant species from 194 flora mapping units are published in this study, but all of them are listed in an electronic appendix. As a result of local surveys, more than 100 new species were found in a single flora mapping unit. Other “linear surveys” along the National Blue Trail revealed 142 new species in 10 flora mapping units in total. Among these, Carex strigosa is one of the most interesting taxa in the Bakony Mts. Previously published and new distribution data of this legally protected sedge species suggests that it is spreading from SW to NE within the Transdanubian Mts. Results from our survey shows that studying synanthropic habitats like frequently used hiking trails might provide a significant amount of new data points at the scale of Hungarian Flora mapping units. Moreover, similar studies might shed some light on the origins and migration routes (e.g. Illyrian and Dacian routes) of “native” species.
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Distribution and identification key of Rubus L. ser. Micantes in Hungary
56-68Views:148Based on a substantial field-work and herbarium revision carried out between 2007 and 2014, the authors present the taxonomical evaluation of recently known representatives of Rubus L. ser. Micantes Sudre in Hungary, providing the first grid-based distribution maps and an identification key of four species of the section (Rubus balatonicus Borbás, R. clusii Borbás, R. styriacus Halácsy, R. tabanimontanus Figert) for Hungary.
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Contributions to the Atlas Florae Hungariae VI.
87-102Views:567The current paper is the sixth one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 389 vascular plant taxa from 117 flora mapping quadrates (CEU) are provided. New records are indicated from different regions of Hungary; however, most occurrences are located in the North Hungarian Mts and from the city of Szolnok and Western Hungary. Occurrence data of rare native taxa (including data of Equisetum fluviatile, Galium rivale, Gladiolus imbricatus, Hypericum maculatum, Persicaria bistorta, Potentilla inclinata, Ranunculus lingua, Rhinanthus rumelicus, Trifolium fragiferum subsp. bonannii, Viola pumila), rare or data-deficient alien taxa (e.g. Amaranthus blitum subsp. blitum, Cymbalaria muralis, Elymus elongatus, Euphorbia peplus, Heracleum sosnowskyi, Impatiens balfourii, Lepidium densiflorum, Rhus typhina, Telekia speciosa), as well as frequent but more or less underrepresented taxa (e.g. Carex otrubae, Chenopodium polyspermum, Crepis tectorum, Filipendula vulgaris, Petrorhagia prolifera, Poa palustris) are also enumerated. In case of the most common species we provided the CEU codes only.
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Contributions to the Atlas Florae Hungariae II.
227-252Views:753The present article is the second part of the series aiming to contribute with new data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 612 vascular plant taxa from 283 flora mapping quadrates (CEU) are presented. New records are distributed over the entire Hungary, however, most are localised in the North Hungarian Mts. Occurrence data of rare native taxa (e.g. Antennaria dioica, Bupleurum pachnospermum, Carex rostrata, Ceratocephala testiculata, Cicuta virosa, Cirsium boujartii, Epipactis moravica, Lycopsis arvensis, Parnassia palustris, Pyrus nivalis, Rumex kerneri, Rumex pulcher, Silene nemoralis, Stellaria alsine, Teucrium botrys, Vicia lutea), rare or data-deficient alien taxa (e.g. Nonea lutea, Panicum dichotomiflorum, Silybum marianum) as well as frequent but more or less underrepresented taxa (e.g. Eragrostis minor, Heliotropium europaeum, Saxifraga tridactylites) are also enumerated, since our intention was to fill the gaps in the Atlas. Remarcable contribution on the occupied territory of Ranunculus illyricus and Spergula pentandra in the Nyírség region (E Hungary), and on the Hungarian distribution of expanding alien weed Senecio vernalis are also presented.
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Contributions to knowledge on the distribution of nine adventive or invasive algae species in Hungary
11-21Views:216Hungarian records of nine adventive or invasive algae [Didymosphenia geminata, Nitzschia closterium, Reimeria sinuata, Navicula schroeteri, Pleurosira laevis (Bacillariophyceae); Pediastrum simplex (Chlorophyceae), Cylindrospermopsis raciborskii, Cuspidothrix issatschenkoi, Sphaerospermopsis aphanizomenoides (Cyanobacteria)] are presented. Navicula schroeteri Meister is a new species for the Hungarian flora. Although the observed distribution patterns of these species were very different, the increasing frequency of their appearance indicates significant effect of climate change on abiotic environment of inland waters, such as the increase in water temperature, increase in salt concentration, and eutrophication. The significance of continuous biomonitoring is emphasized, which, through the recognition of natural and human-induced changes in species composition of algae, or the appearance and rapid expansion of non-indigenous species, can detect environmental changes of inland waters.
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Online distribution atlas of the Hungarian vascular flora (Atlas Florae Hungariae): Homepage structure and prospects of database building
238–252Views:791In December 2018 the online Distribution atlas of vascular plants of Hungary (Atlas Florae Hungariae) was published. The database was built from more than 1 million data records. It mainly contains data from the Hungarian Flora Mapping Programme, but herbarium and literature data were also processed. Maps of species richness illustrate the taxonomic diversity of biogeographical areas and shows the differences of survey quality. After the Flora Mapping Programme ended, between 2016 and 2019, additional data from scientific articles were processed. 25% of new data originated from the article series “Contributions to the Atlas Florae Hungariae” in the Kitaibelia journal, which significantly broadened the distribution of some species. Records of Mesophile forest species (e.g. Allium ursinum, Dryopteris dilatata) increased, as well as records of weed species (e.g. Euphorbia maculata, Senecio vernalis) which can be attributed to their natural spreading. The most important future tasks are to create a reviewed list of taxa and extend data records with an attribute table containing all additional information. Until now the database website is presented only in Hungarian (http://floraatlasz.uni-sopron.hu). Registration is available for anybody, after sending a request via e-mail (atlas.florae.hungariae@uni-sopron.hu). There are different options (excel file by e-mail or online web form) on how to upload floristic data for registered users. Data is only accepted if required information (name of taxa, date, data publisher, settlement, CEU code) is filled out. Each record of uploaded data is revised by the website administrator and taxa specialists. The English version of the webpage will be available in the near future.
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Research on the distribution of woolly cup grass (Eriochloa villosa) in the northern part of Borsod-Abaúj-Zemplén county, Hungary
27–32Views:669Eriochloa villosa (Thunb.) Kunth was recorded for the first time in Hungary in 2007. Since then, the species spread intensively in the area and appeared at some other parts of Hungary. We carried out a research at the current distribution range of the species in the northern part of Borsod-Abaúj-Zemplén county. We asked how big is the infested area surrounding the place of the first occurrence and whether other occurrences could be found in other parts of the investigated area. This article also gives a review on its known localities in Hungary and neighbouring areas. During the field research, 58 locations of the species were identified, it occurred mainly in ploughed and unploughed agricultural fields, and at some places in disturbed grasslands, too. It has spread on a large scale around the neighbouring areas of the first location found in 2007. Other locations were also found even at more distant places such as in the valley of the Sajó and Bódva rivers and in the Cserehát region. This means that further spread of Eriochloa villosa can be expected, especially in areas with an intensive agricultural activity, in disturbed grasslands, and a threat of its appearance is also suspected in open natural habitats.
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Distribution and flora of areas with alkaline and saline soils in the Mezőföld, Hungary
39-61Views:360Areas with alkaline and saline (solonetz and solonchak) soils occupy substantial areas in the westernmost, Transdanubian part of the Great Hungarian Plain (Mezőföld). To this date, the flora and vegetation of only two localities have been studied in some detail, while the rest has remained largely unknown in this respect. In this paper, I provide brief descriptions of these little-known areas and present a list of halophytes with distribution data from the last 17 years. The names of the plants are followed by the name of the township and the geographical location where the plant was found, the code of the grid cell of the Central European Flora Survey grid that includes the location, and in selected species, the year when the species was first recorded. The list is arranged in alphabetical order irrespective of taxonomy. The occurrence data indicate that the floras of these saline-alkaline areas represent only subsets of, but in combination are almost identical to the overall flora of similar habitats in the Sárvíz plain.
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Leucojum vernum at the Súri-Bakonyalja and overview of its distribution in the Bakony Region
242-246Views:568In Hungary Leucojum vernum is considered a plant species of montane character, distributed predominantly in Transdanubia. Apart from some isolated localities (Zalaszántó Basin Mts, Tapolca Basin), the occurrences of the species are mainly confined to the region “Magas Bakony” and its periphery within the Hungarian Middle Mts. In the spring of 2020 a large population was discovered by the author in the region “Súri-Bakonyalja”, which significantly expands the known local distribution range of the species. The new population, together with other phytogeographically significant species (e.g. Anemone nemorosa, Corydalis intermedia, Veratrum album), was found in a poorly explored valley system (Kisbér village: Báró-erdő). Based on all literature and herbarium records, the local distribution range of L. vernum is surveyed.
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New occurrences of Ranunculus strigulosus in East Hungary
3–8Views:591The exact distribution of Ranunculus strigulosus Schur is poorly known in Hungary. Current occurrence data of the species mainly from Békés county and only one archive data from the Nyírség region, from the 1920s were known. This study presents eleven newly discovered occurrences in East Hungary. Populations were found in wet meadows, forest glades and poplar plantations. Nine of these populations are located at the southeastern edge of the Nyírség region and most of them are quite small. Three larger population were found in Nyírábrány, Álmosd and the Tócó Valley between Debrecen and Hajdúböszörmény. One of the newly discovered populations provides a new contribution to the flora of Bátorliget Marsh and Fen-wood NCA. Some morphological characters (hairs on the stem and the leaves, shape of the leaves developed before and after the mowing) are also discussed.
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Rubus armeniacus Focke, an unnoticed invader in the Hungarian flora
220-228Views:231The authors report on the occurrence, distribution and invasion potential of Rubus armeniacus, a bramble species of Caucasian origin, in Hungary. The first verified records of the species are from the 1990s. It has established strong populations in ruderal habitats mainly around Budapest and some other cities, whereas its competiveness is rather weak in closed non-lowland forests rich in native bramble species. In terms of conservation, it constitutes a real threat to the vegetation of Pannonian sandy habitats. The results of this study also emphasize the importance of recognising "nonnative cryptic species" for ecology and conservation biology.
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Contribution to the distribution of Epipactis species in the Keszthely Mts
8-14Views:207In this paper I present occurrence data of Epipactis species in the Keszthely Mts which were collected during the last four years. Ten species had literary or herbarium data from the area previously, all of them has been confirmed (Epipactis atrorubens, E. helleborine, E. leptochila, E. microphylla, E. moravica, E. muelleri, E. palustris, E. purpurata, E. nordeniorum, E. voethii) and another five species (Epipactis albensis, E. neglecta, E. peitzii, E. pontica, E. tallosii) furthermore a hybrid (Epipactis ×reinekei) were found newly on the area.
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Distribution and downstream drift of six native aquatic plant species along irrigation canals in Tiszántúl region (Eastern Hungary)
17–30Views:106Before the nineteenth century, a substantial portion of the Hungarian Great Plain consisted of seasonally or permanently waterlogged habitats, which underwent radical transformation following large-scale water regulation interventions in the Tisza Valley. As a result, extensive wetlands were converted into arable land, causing a major loss of natural vegetation. We surveyed the Eastern (Keleti), Western (Nyugati), and Nagykunság Irrigation Main Canals and selected tributaries to map the distribution of six native wetland species (Carex pseudocyperus, Nymphaea alba, Nymphoides peltata, Trapa natans, Salvinia natans, and Stratiotes aloides). Field surveys were conducted in summer 2022 along the full length of the main canals using 50-m transects spaced ~300–900 m apart (depending on accessibility and vegetation visibility); all sampling locations were recorded in WGS84. To evaluate the potential contribution of water flow to dispersal, we (i) counted drifting individuals of three predominantly free-floating species (S. natans, T. natans and S. aloides) during 60-min observation periods at 26 sites, and (ii) measured surface drift velocity at 24 bridges using 20-cm Phragmites stem fragments (10 replicates per site). All six target species occurred along the Western Main Canal, five along the Eastern Main Canal and three along the Nagykunság Canal. Salvinia natans was the most widespread species, whereas Nymphaea alba was recorded only rarely; Nymphoides peltata was detected exclusively along the Western Main Canal. Surface drift velocity differed markedly among canals, with the highest mean values in the Eastern Main Canal (0.34 ± 0.18 m/s) compared with the Western (0.17 ± 0.03 m/s) and Nagykunság (0.15 ± 0.03 m/s) canals. During drift counts, S. natans was observed at all sites (often >1,000 individuals per hour), T. natans at eight sites (118 individuals in total), and S. aloides as a single drifting specimen. Based on measured drift speeds, a small free-floating macrophyte could be transported up to ~29 km per day in the Eastern Main Canal; order-of-magnitude extrapolations from hourly counts suggest that millions of individuals, particularly S. natans, may be dispersed downstream during the summer vegetation period. Our results indicate that irrigation canals can support native aquatic plant populations and may facilitate downstream dispersal in the Great Plain, highlighting their potential conservation value in an otherwise habitat-poor agricultural landscape.
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Contributions to the Atlas Florae Hungariae XI.: Data from the Tokaj–Zemplén Mts and its surroundings
239–241Views:338This paper is the 11th in the series aiming to contribute data to the distribution maps published recently in Atlas Florae Hungariae. Our records were collected from the Tokaj-Zemplén Mts and the Bodrogköz region. The dataset includes 325 occurrence data of 75 vascular plant taxa from 53 flora mapping unit. Most of the represented taxa are legally protected, e.g. Aconitum variegatum, Alchemilla subcrenata, Aster amellus, Cardamine glanduligera, Centaurea triumfettii, Cephalanthera damasonium, Dianthus deltoides, Epipactis pontica, Gentianopsis ciliata, Orchis morio, Orthilia secunda, Petasites albus, Platanthera bifolia, Polygala major, Prunella grandiflora, Pulsatilla grandis, Rosa pendulina, Salix aurita, Scilla kladnii and Stipa dasyphylla.
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The Himantoglossum adriaticum Baumann in the East-Zala hills (W Hungary)
297–303Views:214This paper presents five new localities of the rare and threatened orchid Himantoglossum adriaticum in Zala county. The new localities falls within the known distribution area of the species. The population size on newly found locations are extremely small, in most cases only 1 generative individual were observed in the mesophilic secondary grasslands of abandoned vineyards. Highly possible, that this species shows a slow, but detectable expansion in Hungary, maybe due to recent climate change process.
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Contributions to the distribution of Arabis nemorensis (Cruciferae) in Hungary
155-169Views:331Arabis nemorensis is a poorly-known species with sparse and mostly uncertain literature records in Hungary. A complete list of its known occurrences was presented in Soó’s synopsis in 1968. A few new records of this taxon have been reported since then. In the present study all literature records were evaluated by means of revision of Hungarian herbarium materials. Several vouchers were succesfully traced, and their identity as A. nemorensis was confirmed (Baja, Ercsi, Lesenceistvánd, Miskolc, Nagykanizsa, Szigetújfalu). In other cases, the putative vouchers belonged to A. hirsuta or A. sagitatta, thus the corresponding literature records proved to be erroneous (Balatonmáriafürdő-alsó, Gyenesdiás, Kőszeg, Vasboldogasszony, Zalaújlak). On the other hand, hitherto unknown Hungarian localities of A. nemorensis were revealed, specifically in the Bükk Mts. (Eger, Kács), the Bakony Mts. (Márkó), in the western Balaton region (Hahót, Hévíz, Keszthely-Fenékpuszta, Nyirád, Tapolca) and the Hungarian Plains (Csákvár, Debrecen, Egeralja, Fertőd-Eszterháza, Kiskőrös, Lébény, Ócsa, Sárszentmihály, Szigetszentmiklós). The species appeared to be new for the phytogeographical regions Bakonyicum and Nyírségense. Additionally, a few specimens of A. nemorensis collected outside the territory of present-day Hungary were revised in BP. The historical occurrence of the species at Torja (Turia in Romania) is documented by two specimens (Schur, 1853, as A. gerardi; Jávorka & Keller, 1943, as A. hirsuta). Another specimen that was collected in 1879 at Óbecse (Bečej) and labelled as A. glastifolia by Kovács was found to be A. nemorensis too. Since the identity of Schneller’s literature record (1858) from Futak is uncertain, Kovács’s specimen is currently the only, though historical record of this species from Vojvodina province in Serbia. Further specimens of A. nemorensis were discovered in the collections of Lengyel. They were collected and labelled as A. sagittata near Pomogy (Pamhagen) in 1910 (and questionably in 1919), probably representing the first gathering of A. nemorensis from Burgenland state in Austria.
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Critical review of the distribution of Equisetum × moorei and E. hyemale in the Nyírség (East Hungary)
131–144Views:664Confusing Moore's horsetail (Equisetum × moorei Newman) with rough horsetail (Equisetum hyemale L.) is a known problem in many countries. In our paper, we review some of the features used for their identification concerning their usability. We examined Equisetum hyemale stands in the Nyírség reported in the literature and E. hyemale specimens of the Herbarium of the University of Debrecen that were collected in the Nyírség. We found that most of these stands and herbarium specimens are actually E. × moorei. The occurrence of E. hyemale was only confirmed in Bátorliget. We also report several new occurrences of Equisetum × moorei in the Nyírség. To better separate the two taxa, we recommend micromorphological examination of the surface of the shoots, which can be performed in the field, as well as observing the height of the leaf-sheath.