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• The herbarium of the Botanical Garden of Eötvös Loránd University (BPU)

  55–59

  Krisztina Nótári

  Timea Nagy

  Viktor Löki

  Tibor Ljubka

  Attila Molnár V.

  Attila Takács

  Views:

  259

  The paper introduces the herbarium of Eötvös Loránd University (BPU), currently stored at the Botanical Garden of the University, according to its state in the year 2013. The BPU herbarium consists of ca. 16 000 specimens the relevant data of which have been organised into an OpenOffice Calc spreadsheet database. The specimens were collected in 16 European countries, mainly in the current territory of Hungary (80%), Romania (12%) and Slovakia (5%). Other countries are represented with very few specimens. The vast majority (ca. 70%) of the Hungarian specimens were collected in Pest, Veszprém, Borsod-Abaúj-Zemplén, Hajdú-Bihar and Szabolcs-Szatmár-Bereg counties. Although the gatherings cover a century and a half period of time, the most rapid growth of the collection took place from the 1930s to the 1960s. The most prolific collectors were Rezső Soó and his followers: Lajos Felföldy, Tibor Simon and Szaniszló Priszter. Felföldy and Simon enriched the collection through more than 60 years. Data on the native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).

• The herbarium of Debrecen University (DE) I. The “Rezső Soó Herbarium”

  142-155

  Attila Takács

  Timea Nagy

  Réka Fekete

  Ádám Lovas-Kiss

  Tibor Ljubka

  Viktor Löki

  Zsuzsa Lisztes-Szabó

  Attila Molnár V.

  Views:

  218

  The paper introduces the herbarium of Debrecen University (DE). All specimens of the herbarium were documented by digital photographs, and all data from the labels were entered into an Excel spreadsheet. The largest collection of Herbarium DE, the “Soó Rezső Herbarium” (abbreviation: SRH) consists of ca. 38.000 specimens from almost all European countries, but mainly from Hungary and the neighbouring countries. Most of the specimens originate from Hajdú-Bihar, Pest, Győr-Moson-Sopron, Borsod-Abaúj-Zemplén, Veszprém and Szabolcs-Szatmár-Bereg counties. The SRH stores a considerable amount of original/authentic specimens collected by Hungarian botanists, primarily by Rezső Soó and his followers, as well as Sándor Polgár. The most rapid growth of the SRH took place in the first half of the last century (1930–1950). By contrast, the collection was practically not developed between 1955 and 2000. After the millennium it started to grow again. Data on the most substantial part of the SRH covering native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting date, collector, file name of documentary photograph).

• Remnants of closed oak woods on loess in the Mezőföld (Pulmonario mollis-Quercetum roboris Kevey 2008)

  66–93

  Balázs Kevey

  András Horváth

  Gábor Lendvai

  Views:

  185

  The phytosociological characterization of closed oak forests occurring in the loess-covered part of Central Hungary (Mezőföld) is presented. Such forest fragments are found very sporadically in that predominantly treeless region, mostly in semi-dry habitats of north-facing hillsides. This plant community represents an intermediate stage between open steppe woodland (Aceri tatarici-Quercetum pubescentis-roboris) and closed mesic oak-hornbeam forests (Corydali cavae-Carpinetum). Analyses of 20 phyto­sociological samples using multivariate grouping methods (cluster analysis, principal coordinates analysis) showed a clear difference between these samples and samples of similar communities occurring in the area. In the species composition the characteristic elements of dry and mesic oak forests (Quercetea pubescentis-petraeae, Quercetalia cerridis, Aceri tatarici-Quercion and Fagetalia, respectively) played a major role. On the other hand, dry grassland species (Festuco-Brometea, Festucetalia valesiacae, Festucion rupicolae, etc.) were much less significant than in steppe woodlands. In terms of chorology, European and sub-Mediterranean floristic elements were dominant in the samples, while the proportion of continental elements was substantially smaller than that in steppe woodlands. As a result, our samples were identified with the plant association Pulmonario mollis-Quercetum roboris Kevey 2008, which is classified into the suballiance Polygonato latifolio-Quercenion roboris Kevey 2008 in the phytosociological system.

• Oak-ash-elm forests in the Nyírség

  179-220

  Balázs Kevey

  László Papp

  Gábor Lendvai

  Views:

  155

  This paper presents phytosociological analyses of oak-ash-elm forests of Nyírség, NE Hungary. These climatically azonal forests grow in habitats where the soil is under the continual influence of the groundwater table, which maintains rather mesic conditions. They are particularly rich in Fagetalia elements (Actaea spicata, Allium ursinum, Anemone ranunculoides, Asarum europaeum, Athyrium filix-femina, Cardamine bulbifera, Carex pilosa, Cerastium sylvaticum, Chaerophyllum aromaticum, Corydalis cava, Dryopteris filix-mas, Epipactis helleborine agg., Euphorbia amygdaloides, Galeobdolon luteum, Galium odoratum, Hedera helix, Isopyrum thalictroides, Lathraea squamaria, Lathyrus vernus, Lilium martagon, Majanthemum bifolium, Mercurialis perennis, Milium effusum, Polygonatum multiflorum, Pulmonaria officinalis, Ranunculus cassubicus, Salvia glutinosa, Sanicula europaea, Scilla vindobonensis, Stachys sylvatica, Stellaria holostea etc.) typical in the submontane regions of this part of Europe. It is possible that they are remnants of the vegetation dominating the region under the less continental and more moist climate during the Beech phase (2500–800 BC) of the Holocene.

• Contributions to the flora of Baranya and Tolna counties III.

  39-50

  István Zsolt Tóth

  Views:

  149

  During the last four years (2014–2017) that have passed since my last publication, I have been collecting 25 000 new floristic data for the territory of the Danube–Drava National Park. The most valuable and interesting records of 116 taxa are presented here, e.g. those of the strictly protected Digitalis ferruginea and Paeonia banatica, and the protected species Polystichum lonchitis, Dryopteris affinis, Doronicum orientale, Helleborus dumetorum, Corydalis intermedia, Spiraea media, Veratrum nigrum. New records of not protected but rare species, such as Lycopsis arvensis, Silene dichotoma, Teucrium botrys, Thladiantha dubia, Vicia lutea, Hypochoeris radicata are also enumerated. Some curiosities are mentioned as well, including the sizes of the biggest (and probably the oldest) Crataegus monogyna, Hedera helix, Ruscus hypoglossum and Ruscus aculeatus individuals that were found during my field work.

• New montane species in the Hungarian bryophyte flora

  16-26

  Csaba Németh

  András Schmotzer

  Views:

  681

  During a bryofloristical investigation in the vicinity of Ómassa (Miskolc) in the Bükk Mountains (NE Hungary), three montane-boreal bryophyte species new to the Hungarian bryoflora were collected. Encalypta spathulata Müll.Hal. was found on the ledges of limestone cliffs of Vörös-kő Hill as well as on the rocky debris among roots of a fallen tree lying under the cliffs. Hydrogonium croceum (Brid.) Jan Kučera and Orthothecium rufescens (Dicks. ex Brid.) Schimp. were found in the crevices of dolomite cliffs on the north-facing steep slope of Jávor Hill. While the latter two species are widespread in European Mountains, E. spathulata is considered to be a rare species, and is assessed as ‘Vulnerable’ in the IUCN red list.

• Contributions to the flora of Budapest and its surroundings II.

  33–50

  Lajos Somlyay

  Sándor Makádi

  Miklós Csábi

  Views:

  230

  New and recently confirmed localities of 52 rare taxa (including 26 orchids) are presented from the vicinity of Budapest. Epipactis peitzii is a new species for the Hungarian flora; it has been recorded in the Buda and Pilis Mts. This species and its closest relatives (E. leptochila group) are briefly discussed. Other records are new for the flora of particular regions, specifically: Buda Mts: Corydalis intermedia, Epipactis leptochila, Epipactis muelleri; Pilis Mts: Anacamptis coriophora, Epipactis neglecta, E. tallosii, Valerianella pumila; Visegrád Mts: Epipactis futakii, E. leptochila, E. muelleri, E. neglecta, Ophrys apifera, O. holubyana.

• A new alien species in Hungary: Limnobium laevigatum (Hydrocharitaceae)

  9-15

  Norbert Riezing

  Views:

  262

  The South American Spongeplant (Limnobium laevigatum (Humb. & Bonpl. ex. Willd.) Heine) is a floating aquatic plant native to freshwater habitats of tropical and subtropical Central and South America. It is frequently used for ornamental purposes in ponds and aquariums, and became invasive in the recent decades around the world. It has a high reproductive potential and a high dispersal capacity as well. It can form massive floating mats causing light limitation and creating anoxic conditions in the underlying water column, which strongly reduces native animal and plant biomass and diversity. It can also hamper navigation and water flow in rivers and canals. It has been introduced to the United States, Australia, Indonesia, Japan, Zambia and Zimbabwe. There is only one previous record in Europe (Belgium). This article is about the first record of Limnobium laevigatum in Hungary. I found two localities situated near Tata-Naszály and Dunaalmás (North-western part of Hungary). Both localities are fed by hot-water springs. It forms a small but dense population in Dunaalmás located near to the hot spring. The population in Tata-Naszály can be found in a 1.3 km long section of a stream, where it formed a sparse population in 2018.

• The Wildflower of the Year 2014 in Hungary: Siberian flag (Iris sibirica L.)

  268-285

  Attila Takács

  Timea Nagy

  Éva Salamon-Albert

  Attila Molnár V.

  Views:

  190

  In this paper a review of the nomenclature, etymology, taxonomy, morphology, histology, life cycle, phenology, reproduction, habitat preference, biotic interactions, biologically active compounds, micropropagation, application possibilities and conservation status of Siberian flag (Iris sibirica L.) can be found. Leaf traits, phenological data, seed-set, thousand-seed weight, germination, growth rate and soil characteristic data are published based on original observations:

  • Leaf area is between 25,3 and 52,9 cm², its dry mass is 232 and 272 mg/g, specific leaf area is 14,5 and 15,0 m²/kg; based on measurement of 5–5 leaves of sibirica, collected from Tapolca and Létavértes (Hungary) in May of 2014.
  • Based on herbarium dataset, blooming of sibirica begins at the end of April and lasts to early-July, contrary to the literature data (May–June).
  • Capsules contain (0–)58–76(–121) fertile seeds. (20–)60–80(–90)% of ovules develops to (seemingly) viable seeds, meanwhile the other ovules remain as aborted ones or develop to infertile seeds (probably because of absence of resources); based on fruits collected from Regéc (Hungary) in 2014 and 2015.
  • Thousand-seed weight of sibirica is 8,8298–11,2914 g (based on 3×100–100 seeds collected from Regéc and Tapolca in 2014 and 2015), which is lower value than the literature data.
  • In our germination test (50–50 seeds sowed to wet soil, after different treatments) 14% of scarified seeds, 6% of scalded seeds, 4–4% of imbibed and control seeds, 0–0% of cooled and refrigerated seeds are germinated. Scarified seeds germinated in the 8–26^th days, imbibed seeds 15–19^th days, control seeds 16–20^th days, scalded seeds 20–23^th days after sowing. It seems that scarification stimulates, meanwhile temperature-treatments inhibit the germination. Until 18 weeks the seedlings grow to 30 cm (in mean) and develop 5–7 leaves (in mean). After the 14^th week, the first and littlemost lateral leaves are necrosed.
  • Analyses of soil samples collected from 17 locations of sibirica in Hungary, suggest that the species prefers highly acidic to slightly alcalic, lime-free to highly calcareous soils with generally high amount of humus and clay, different amount of phosphorus, potassium and nitrogen and low concentration of salt.

• Floristic data from Lengyel–Hőgyész Natura 2000 site (SW Hungary)

  31–38

  István Vas

  István Zsolt Tóth

  Views:

  168

  This paper presents occurrence data of 169 taxa from the Lengyel–Hőgyész Natura 2000 site (HUDD20026) which is on the southern part of Tolna Hills, SW Hungary. Data were collected during three years (2015, 2016, 2017), on different sites annually. The presented data contribute to the distribution dataset of Atlas florae Hungariae. Regionally rare or sporadic species (e.g. Asplenium scolopendrium, Asplenium trichomanes, Polystichum setiferum, Doronicum hungaricum, Paris quadrifolia, Scilla vindobonensis) are enumerated. Presence of Epipactis leptochila subsp. neglecta and Platanthera chlorantha were not registered from this site before. Subspecies of Carex divulsa are also studied on this site – distribution of these infraspecific taxa have not been mapped recently in Hungary.

• Data to the flora of Hungary I.

  254-259

  Dániel Kovács

  Views:

  80

  The paper gives new floristic data from several regions of Hungary. Some of them are recent evidences of old data (e.g. Herniaria incana in Pomáz), others are new occurrences of recently expanding alien species (e.g. Eleusine indica, Geranium purpureum, Typha laxmannii), casual species (e.g. Lobularia maritima, Nepeta racemosa) or legally protected taxa (e.g. Agrostemma githago, Ornithogalum brevistylum, Ranunculus illyricus, Sonchus palustris).

• The purple viper's bugloss (Echium plantagineum), a new adventive species for Hungary

  199-206

  Anikó Csecserits

  Gusztáv Jakab

  Tamás Rédei

  Views:

  898

  Following global trends, the number of newly established alien plant species is increasing in Hungary. Not only professional scientists but also citizens could contribute to the discovery and documentation of new occurrences. One of the first records of the purple viper’s bugloss (Echium plantagineum L.) in Hungary originate from an online group dealing with wild plant species identification. This species is native in Western Mediterranean countries, and is introduced to other continents. In Australia, it is a well-spread, largely problematic invasive species, and it might show similar tendencies in Hungary as well. In order to acquaint the wider public with this species, we present its morphology, phenology and other characteristics and provide a possible inlay in the Hungarian identification key.

• Epipactis pseudopurpurata Mered’a, a new orchid species for the Hungarian flora

  27–32

  Miklós Csábi

  Antal Halász

  Views:

  113

  A small population of Epipactis pseudopurpurata new to the flora of Hungary was found in the Buda Mts, near Telki village (Pest county). The main diagnostic features of this species are discussed, and a comparison with the allied species, E. purpurata is provided.

• Distribution and identification key of Rubus L. ser. Micantes in Hungary

  56-68

  Gergely Király

  Bohumil Trávnícek

  Vojtěch Žíla

  Views:

  83

  Based on a substantial field-work and herbarium revision carried out between 2007 and 2014, the authors present the taxonomical evaluation of recently known representatives of Rubus L. ser. Micantes Sudre in Hungary, providing the first grid-based distribution maps and an identification key of four species of the section (Rubus balatonicus Borbás, R. clusii Borbás, R. styriacus Halácsy, R. tabanimontanus Figert) for Hungary.

• The occurrence of Viola collina Besser in the Bükk and Uppony Mts. (N Hungary)

  254-258

  József Sulyok

  András Vojtkó

  Lajos Somlyay

  Views:

  159

  Herbarium revisions and recent field observations revealed the occurrence of Viola collina Besser in the Bükk and Uppony Mts. (northern Hungary). This species has not been reported from the North Hungarian Mountains thus far. Altogether 94 localities of V. collina have been recorded in that region between 2007 and 2014. A map of the species’ local distribution is provided. Local habitat preferences of V. collina are briefly discussed.

• Historical occurrence of Spergularia echinosperma, a rare Nanocyperion species in Hungary

  3–9

  Attila Mesterházy

  Pavel Kúr

  Views:

  300

  Three species of the genus Spergularia have been known from Hungary. The most widespread one, Spergularia rubra (L.) J. Presl et C. Presl occurs on young sandy fallow fields and other disturbed habitats.  The other species, S. marina (L.) Bess. and S. media (L.) C. Presl, occur sporadically in saline grasslands, and both have been recently spreading along salted roads. During a revision of the genus Spergularia in Hungarian herbaria, specimens of the species S. echinosperma (Čelak.) Asch. et Graebn. were found. It is a rare species previously known to occur only in Czechia and Germany and to be extinct in Austria and Slovakia (a few occurrences along the Morava and Danube rivers between 1909 and 1953). However, it was recently discovered in Western Siberia and Kazakhstan. It is a typical mudflat plant that occurs on exposed sandy shores of water bodies. Two herbarium sheets collected by Gyula Tauscher from near the village of Ercsi (riverbank of the Danube) in 1903 were found in the Carpato-Pannonicum collection of the Hungarian Natural History Museum. In addition, a new specimen was found, collected in Slovakia (Danube near Šamorín, collected by Mihály Résely in 1868) at the vicinity of the Hungarian border. The species was not confirmed during a field inspection of the target localities in Hungary; therefore, we have to consider it extinct in the country as well as in the whole Pannonian Basin. Recently, S. echinosperma was split into two subspecies – subsp. echinosperma and subsp. albensis. The subsp. echinosperma is probably restricted to fishponds in the SW part of Czechia and subsp. albensis occurs mainly in the floodplains of the river Elbe in Germany. The extinct populations of S. echinosperma in the Pannonian Basin belonged to the subsp. albensis. This taxon is morphologically close to S. rubra but can be reliably distinguished by densely verrucose seeds and shorter than wide stipules.

• Some floristic data from surroundings of Kötcse II. (West Outer Somogy, South Transdanubia, Hungary)

  114-121

  Timea Nagy

  Views:

  113

  Floristic data of 73 species from NE part of West Outer Somogy are provided in the paper. Most of them were collected in 2016. The paper contains distribution data of species, which have not been detected in Outer Somogy so far (e.g. Peucedanum arenarium), and new occurrences of rare or sporadic taxa (e.g. Menyanthes trifoliata, Veronica scardica), as well as subspontaneous occurrences of cultivated species (e.g. Narcissus poëticus, Helleborus purpurascens) are provided.

• Cyperus odoratus in Hungary

  157–164

  Attila Mesterházy

  Views:

  331

  Eleven species of the genus Cyperus have been reported from Hungary so far, of which 3 are certainly aliens. A small population of C. odoratus, new for the flora of Hungary, was found along the River Danube near Foktő settlement (Bács-Kiskun County, S Hungary) in late autumn of 2019. On this occasion blooming specimens were observed in the floodplain ruderal vegetation on the bank of a fork of the Danube. This species is widespread in the tropical and subtropical regions of the World, and was first introduced to Europe in the early 1950s.  More recently it has been spreading along certain rivers of Italy and Spain. It was discovered along the river Danube in Romania in the early 1990s, and later at the upper parts of the river in Bulgaria, Croatia and Serbia. Its introduction into Hungary is connected with the Danube, too. Based on foreign experience, it is probable that C. odoratus will be spreading along the Hungarian Danube system, hopefully, not as an invasive species.

• Contributions to the lichen flora of the Mecsek Mts (South Hungary)

  16–26

  Dániel Kovács

  László Lőkös

  Views:

  98

  A checklist of 115 lichen taxa was compiled from the Mecsek Mts, based on floristical records of 221 old or recently collected herbarium specimens (incl. 37 duplicates) deposited in the JPU herbarium (Pécs, Hungary). The occurrence of 22 taxa are new for the Mecsek Mts: Arthopyrenia cinereopruinosa, Baeomyces rufus, Candelariella reflexa, Catapyrenium rufescens, Cladonia macilenta, Dimerella pineti, Hypogymnia tubulosa, Ochrolechia arborea, Pleurosticta acetabulum, Pseudevernia furfuracea, Psilolechia lucida, Pycnothelia papillaria, Rinodina sophodes, Tomasellia arthonioides, Toninia candida, Trapelia involuta, Varicellaria hemisphaerica, Verrucaria marmorea, V. nigrescens, V. parmigera, Xanthoparmelia conspersa, X. verruculifera. One legally protected lichen species (Peltigera leucophlebia) is also represented by two specimens.

• Floristic data from the Great Plain of Hungary (Alföld)

  304–316

  Márton Korda

  Dávid Schmidt

  Róbert Vidéki

  Győző Haszonits

  Viktor Tiborcz

  Ágnes Csiszár

  Gergely Zagyvai

  Dénes Bartha

  Views:

  230

  In this study we report relevant occurrence data of 49 species and hybrids and their habitats from the Great Plain registered in the period between 2011 and 2017. Due to importance for nature conservation, we present additional data of further 13 species. Most of the data are originated from riverine oak-elm-ash forests of South East Hungary (from the region of Gyula, Békéscsaba, Doboz settlements). The most important result of our study is the rediscovery of Gagea minima and Dictamnus albus. Besides, we report new occurrence data for the Great Plain (Alföld), as well as for the flora of the microregions in questions (e.g. Draba muralis, Nicandra physalodes, Cystopteris fragilis) and also present the data of the regionally endangered species (e.g. Inula helenium, Ophioglossum vulgatum). Some rare weed taxa (e.g. Calepina irregularis, Cardamine impatiens), spreading adventive taxa (e.g. Chorispora tenella, Euphorbia maculata, Phytolacca americana, Ph. esculenta, Sicyos angulata) and in surveys rather underrepresented species (e.g. Loranthus europaeus, Arabis hirsuta, Rumex confertus) are reported too.

• The occurences of two ‘old-new’ alien grass species, Rostraria cristata (L.) Tzvelev and Cynosurus echinatus L. in Pécs

  39-42

  Tamás Wirth

  Attila Lengyel

  Views:

  91

  Some small populations of Rostraria cristata and Cynosurus echinatus, two ‘old-new’ alien grass species in Hungary, were recently discovered in the city of Pécs (South Hungary). A morphological description of these species as well as a survey of their former and current localities are presented. 

• A new montane bryophyte species from the Mátra Mts, Hungary: Hymenoloma crispulum

  11-18

  Csaba Németh

  Peter Erzberger

  Views:

  237

  Due to the geomorphological situation and climatic conditions of Hungary, true montane climate is restricted to very few and small areas of higher elevation, with the highest peak (Mt Kékes, 1014 m) in the Mátra Mountains (North Central Hungary). During a bryofloristical survey in this re­gion, a few cushions of a Boreo-arctic montane bryophyte species, Hymenoloma crispulum (Hedw.) Ochyra (syn.: Dicranoweisia crispula (Hedw.) Milde), was discovered at the steep northern slope of Mt. Kékes in an andesite boulder scree with cool and humid microclimate. H. crispulum is new to the Hun­garian bryoflora and fits in well with other co-occurring vascular and cryptogamic taxa of northern-montane affinities. The population is very small and vulnerable, therefore it should be rated 'critically endangered' (CR) in an updated national red list.

• Bruchia flexuosa (Schwägr.) Müll. Hal. a true peculiarity in the Hungarian bryoflora

  193-201

  Csaba Németh

  Attila Mesterházy

  Views:

  140

  The fifth European occurrence of the moss species Bruchia flexuosa possessing a holarctic distribution range with a North American centre was encountered in Hungary. Outside of the North American continent only two Asian (both from Japan) and four European occurrences from four different countries (Italy, Austria, Slovenia, Croatia) have hitherto been reported. Apart from the recent Japan findings and the Croatian record observed in 1987, all other European records date back to the 19^th century. Our paper reports the circumstances of the Hungarian occurrence along with a short descriptions and photographic illustrations of both the species and its habitat.

• Escaping of Euphorbia myrsinites from cultivation in Eger (E Hungary)

  253–256

  Peregrym Mykyta

  Views:

  170

  Euphorbia myrsinites L. (Euphorbiaceae) is a well-known ornamental plant with native dis­tribu­tion around the Mediterranean Basin, the Black Sea, the Caucasus region and Minor Asia. Its es­cape from cultivation and its naturalization in Western and Central Europe as well as in North America was reported in many cases. In Hungary, the casual escape of E. myrsinites has been documented more than fifty years ago and new observations were recently reported from different regions of the country. Here, I discuss a new record of the species from the town of Eger (E Hungary). The obtained data allow considering E. myrsinites as a locally naturalized species in the flora of Hungary. According to standard­ized criteria, the species may be able for local invasions too, but further observations are needed to accurately determine its present status in Hungary.

• Data to the flora and vegetation of Hungary II.

  104–113

  Eszter Aradi

  László Erdős

  Viktória Cseh

  Csaba Tölgyesi

  Zoltán Bátori

  Views:

  188

  We report the occurrence data of 45 vascular plant taxa collected at various localities in Hungary (Duna–Tisza Interfluve, Tiszántúl and South Transdanubia) during floristic, vegetation and ecological field surveys mostly between 2013 and 2016. Many of these plants are included in the ’Red list of the vascular flora of Hungary’ (e.g. Astragalus dasyanthus, Erodium ciconium and Lotus angustissimus). We primarily focused on the occurrence of rare plants (e.g. Apium repens, Epipactis bugacensis and Trifolium strictum), plants that appeared in unexpected places (e.g. Medicago arabica, Polypodium vulgare and Polystichum setiferum) or have a phyto-geographic importance (e.g. Piptatherum virescens, Ranunculus psilostachys and Tamus communis).