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  • Contributions to the flora of the Szuha watershed and adjacent territories (NE Hungary)
    27-67
    Views:
    307

    This paper presents new floristic data of 415 taxa from the Putnok Hills and Sajó Valley microregions (Northern Hungary). Records were collected between 1991 and 2021. In the first part new occurrences of legally protected (e.g., Cypripedium calceolus, Iris aphylla subsp. hungarica, Lycopodium clavatum, Thalictrum aquilegiifolium, Pyrola rotundifolia, Scorzonera purpurea, Stipa pulcherrima) and not protected but locally rare or data-deficient species (e.g., Myosurus minimus, Thalictrum simplex, Sisymbrium loeselii, Camelina microcarpa, Potentilla collina, Trifolium striatum, Geranium pratense, G. dissectum, G. divaricatum, Viola pumila, Epilobium roseum, Myriophyllum spicatum, Pimpinella major, Berula erecta, Trinia ramosissima, Centaurium pulchellum, Nepeta cataria, Limosella aquatica, Veronica scutellata, Orobanche arenaria, Utricularia vulgaris, Galium elongatum, Cephalaria transsylvanica, Dipsacus fullonum, Xanthium saccharatum, Alisma lanceolatum, Elodea canadensis, Potamogeton pectinatus, P. berchtoldii, P. lucens, P. nodosus, Zannichellia palustris, Sisyrinchium bermudiana, Melica picta, Elymus elongatus, Leersia oryzoides, Typha laxmannii, Eriochloa villosa) are presented. This chapter contains data on 246 species from which 8 are new for the flora of the Tornense flora region and 47 for its subregion, the Putnok Hills. Further contributions to the distribution data of Atlas Florae Hungariae are presented in Electronic Appendix. This section contains raw occurrence data of 169 species.

  • White willow riparian forests along the upper Tisza River, Hungary
    78–100
    Views:
    131

    Forests of the Tisza floodplain in the northwestern part of Hungary include white willow riparian forests (Leucojo aestivi-Salicetum albae) that are phytosociologically little studied. This study summarizes the characteristics of this community based on 25 phytosociological relevés. These forests grow mostly on mud and raw alluvial soil in the low-lying parts of the lower terraces of the floodplain. They are easily separated by their species composition and underdeveloped shrub layer from white poplar gallery forests (Senecioni sarracenici-Populetum albae)., which grow on higher lying ground with rather light sandy soil and typically possess a pronounced shrub layer. The understory of their stands may often host plants  that are rare or completely absent in other parts of the country, such as Cardamine amara, Cardamine flexuosa, Cardaminopsis arenosa, Carex pseudocyperus, Carex remota, Chrysosplenium alternifolium, Leucanthemella serotina, Leucojum aestivum, Oenanthe banatica, Scrophularia scopolii, Telekia speciosa, Vitis sylvestris. This community is classified in the „Salicenion albae-fragilis Kevey 2008” suballiance.

  • White poplar riparian forests on the Csepel-sziget, Middle-Hungary (Senecioni sarracenici-Populetum albae Kevey in Borhidi & Kevey 1996)
    57–78
    Views:
    224

    In this paper the white poplar riparian forests (Senecioni sarracenici-Populetum albae) growing on the Csepel-sziget and its vicinity are described and characterized based on 25 phytosocio­logical relevés. These communities grow on loose fluvial sand and raw alluvial soils on the elevated parts of the lower river floodplain. They can readily be distinguished from willow gallery forests (Leucojo aestivi-Salicetum albae) which have no shrub layers and grow in habitats 1–1.5 m below the level of poplar forests on rather heavy and muddy soils. They differ also from the oak-ash-elm forests (Scillo vindobonensis-Ulmetum) growing in the upper floodplain. Certain – partly submontane – plants that are rare or completely absent in other parts of the Great Hungarian Plains may also occur in them, such as Anemone ranunculoides, Carex remota, Clematis recta, Crataegus × degenii, Crataegus nigra, Epipactis helleborine, Equisetum hyemale, Galanthus nivalis, Lathraea squamaria, Leucojum aestivum, Paris quadrifolia, Scilla vindobonensis, Vitis sylvestris. This association is classified in the sub-alliance Populenion nigro-albae Kevey 2008 in the syntaxonomical system.

  • The herbarium of Debrecen University (DE) I. The “Rezső Soó Herbarium”
    142-155
    Views:
    216

    The paper introduces the herbarium of Debrecen University (DE). All specimens of the herbarium were documented by digital photographs, and all data from the labels were entered into an Excel spreadsheet. The largest collection of Herbarium DE, the “Soó Rezső Herbarium” (abbreviation: SRH) consists of ca. 38.000 specimens from almost all European countries, but mainly from Hungary and the neighbouring countries. Most of the specimens originate from Hajdú-Bihar, Pest, Győr-Moson-Sopron, Borsod-Abaúj-Zemplén, Veszprém and Szabolcs-Szatmár-Bereg counties. The SRH stores a considerable amount of original/authentic specimens collected by Hungarian botanists, primarily by Rezső Soó and his followers, as well as Sándor Polgár. The most rapid growth of the SRH took place in the first half of the last century (1930–1950). By contrast, the collection was practically not developed between 1955 and 2000. After the millennium it started to grow again. Data on the most substantial part of the SRH covering native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting date, collector, file name of documentary photograph).

  • The herbarium of the Botanical Garden of Eötvös Loránd University (BPU)
    55–59
    Views:
    256

    The paper introduces the herbarium of Eötvös Loránd University (BPU), currently stored at the Botanical Garden of the University, according to its state in the year 2013. The BPU herbarium consists of ca. 16 000 specimens the relevant data of which have been organised into an OpenOffice Calc spreadsheet database. The specimens were collected in 16 European countries, mainly in the current territory of Hungary (80%), Romania (12%) and Slovakia (5%). Other countries are represented with very few specimens. The vast majority (ca. 70%) of the Hungarian specimens were collected in Pest, Veszprém, Borsod-Abaúj-Zemplén, Hajdú-Bihar and Szabolcs-Szatmár-Bereg counties. Although the gatherings cover a century and a half period of time, the most rapid growth of the collection took place from the 1930s to the 1960s. The most prolific collectors were Rezső Soó and his followers: Lajos Felföldy, Tibor Simon and Szaniszló Priszter. Felföldy and Simon enriched the collection through more than 60 years. Data on the native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).

  • Remnants of closed oak woods on loess in the Mezőföld (Pulmonario mollis-Quercetum roboris Kevey 2008)
    66–93
    Views:
    182

    The phytosociological characterization of closed oak forests occurring in the loess-covered part of Central Hungary (Mezőföld) is presented. Such forest fragments are found very sporadically in that predominantly treeless region, mostly in semi-dry habitats of north-facing hillsides. This plant community represents an intermediate stage between open steppe woodland (Aceri tatarici-Quercetum pubescentis-roboris) and closed mesic oak-hornbeam forests (Corydali cavae-Carpinetum). Analyses of 20 phyto­sociological samples using multivariate grouping methods (cluster analysis, principal coordinates analysis) showed a clear difference between these samples and samples of similar communities occurring in the area. In the species composition the characteristic elements of dry and mesic oak forests (Quercetea pubescentis-petraeae, Quercetalia cerridis, Aceri tatarici-Quercion and Fagetalia, respectively) played a major role. On the other hand, dry grassland species (Festuco-Brometea, Festucetalia valesiacae, Festucion rupicolae, etc.) were much less significant than in steppe woodlands. In terms of chorology, European and sub-Mediterranean floristic elements were dominant in the samples, while the proportion of continental elements was substantially smaller than that in steppe woodlands. As a result, our samples were identified with the plant association Pulmonario mollis-Quercetum roboris Kevey 2008, which is classified into the suballiance Polygonato latifolio-Quercenion roboris Kevey 2008 in the phytosociological system.

  • Oak-ash-elm forests in the Nyírség
    179-220
    Views:
    149

    This paper presents phytosociological analyses of oak-ash-elm forests of Nyírség, NE Hungary. These climatically azonal forests grow in habitats where the soil is under the continual influence of the groundwater table, which maintains rather mesic conditions. They are particularly rich in Fagetalia elements (Actaea spicataAllium ursinum, Anemone ranunculoides, Asarum europaeum, Athyrium filix-femina, Cardamine bulbifera, Carex pilosa, Cerastium sylvaticum, Chaerophyllum aromaticum, Corydalis cava, Dryopteris filix-mas, Epipactis helleborine agg., Euphorbia amygdaloides, Galeobdolon luteum, Galium odoratum, Hedera helix, Isopyrum thalictroides, Lathraea squamaria, Lathyrus vernus, Lilium martagon, Majanthemum bifolium, Mercurialis perennis, Milium effusum, Polygonatum multiflorum, Pulmonaria officinalis, Ranunculus cassubicus, Salvia glutinosa, Sanicula europaea, Scilla vindobonensis, Stachys sylvatica, Stellaria holostea etc.) typical in the submontane regions of this part of Europe. It is possible that they are remnants of the vegetation dominating the region under the less continental and more moist climate during the Beech phase (2500–800 BC) of the Holocene.

  • Contributions to the flora of Baranya and Tolna counties III.
    39-50
    Views:
    142

    During the last four years (2014–2017) that have passed since my last publication, I have been collecting 25 000 new floristic data for the territory of the Danube–Drava National Park. The most valuable and interesting records of 116 taxa are presented here, e.g. those of the strictly protected Digitalis ferruginea and Paeonia banatica, and the protected species Polystichum lonchitis, Dryopteris affinis, Doronicum orientale, Helleborus dumetorum, Corydalis intermedia, Spiraea media, Veratrum nigrum. New records of not protected but rare species, such as Lycopsis arvensis, Silene dichotomaTeucrium botrys, Thladiantha dubia, Vicia lutea, Hypochoeris radicata are also enumerated. Some curiosities are mentioned as well, including the sizes of the biggest (and probably the oldest) Crataegus monogyna, Hedera helix, Ruscus hypoglossum and Ruscus aculeatus individuals that were found during my field work.

  • New montane species in the Hungarian bryophyte flora
    16-26
    Views:
    674

    During a bryofloristical investigation in the vicinity of Ómassa (Miskolc) in the Bükk Mountains (NE Hungary), three montane-boreal bryophyte species new to the Hungarian bryoflora were collected. Encalypta spathulata Müll.Hal. was found on the ledges of limestone cliffs of Vörös-kő Hill as well as on the rocky debris among roots of a fallen tree lying under the cliffs. Hydrogonium croceum (Brid.) Jan Kučera and Orthothecium rufescens (Dicks. ex Brid.) Schimp. were found in the crevices of dolomite cliffs on the north-facing steep slope of Jávor Hill. While the latter two species are widespread in European Mountains, E. spathulata is considered to be a rare species, and is assessed as ‘Vulnerable’ in the IUCN red list.

  • Contributions to the flora of Budapest and its surroundings II.
    33–50
    Views:
    227

    New and recently confirmed localities of 52 rare taxa (including 26 orchids) are presented from the vicinity of Budapest. Epipactis peitzii is a new species for the Hungarian flora; it has been recorded in the Buda and Pilis Mts. This species and its closest relatives (E. leptochila group) are briefly discussed. Other records are new for the flora of particular regions, specifically: Buda Mts: Corydalis intermediaEpipactis leptochilaEpipactis muelleri; Pilis Mts: Anacamptis coriophoraEpipactis neglectaE. tallosiiValerianella pumila; Visegrád Mts: Epipactis futakiiE. leptochilaE. muelleriE. neglectaOphrys apiferaO. holubyana.

  • The Wildflower of the Year 2014 in Hungary: Siberian flag (Iris sibirica L.)
    268-285
    Views:
    184

    In this paper a review of the nomenclature, etymology, taxonomy, morphology, histology, life cycle, phenology, reproduction, habitat preference, biotic interactions, biologically active compounds, micropropagation, application possibilities and conservation status of Siberian flag (Iris sibirica L.) can be found. Leaf traits, phenological data, seed-set, thousand-seed weight, germination, growth rate and soil characteristic data are published based on original observations:

    • Leaf area is between 25,3 and 52,9 cm2, its dry mass is 232 and 272 mg/g, specific leaf area is 14,5 and 15,0 m2/kg; based on measurement of 5–5 leaves of sibirica, collected from Tapolca and Létavértes (Hungary) in May of 2014.
    • Based on herbarium dataset, blooming of sibirica begins at the end of April and lasts to early-July, contrary to the literature data (May–June).
    • Capsules contain (0–)58–76(–121) fertile seeds. (20–)60–80(–90)% of ovules develops to (seemingly) viable seeds, meanwhile the other ovules remain as aborted ones or develop to infertile seeds (probably because of absence of resources); based on fruits collected from Regéc (Hungary) in 2014 and 2015.
    • Thousand-seed weight of sibirica is 8,8298–11,2914 g (based on 3×100–100 seeds collected from Regéc and Tapolca in 2014 and 2015), which is lower value than the literature data.
    • In our germination test (50–50 seeds sowed to wet soil, after different treatments) 14% of scarified seeds, 6% of scalded seeds, 4–4% of imbibed and control seeds, 0–0% of cooled and refrigerated seeds are germinated. Scarified seeds germinated in the 8–26th days, imbibed seeds 15–19th days, control seeds 16–20th days, scalded seeds 20–23th days after sowing. It seems that scarification stimulates, meanwhile temperature-treatments inhibit the germination. Until 18 weeks the seedlings grow to 30 cm (in mean) and develop 5–7 leaves (in mean). After the 14th week, the first and littlemost lateral leaves are necrosed.
    • Analyses of soil samples collected from 17 locations of sibirica in Hungary, suggest that the species prefers highly acidic to slightly alcalic, lime-free to highly calcareous soils with generally high amount of humus and clay, different amount of phosphorus, potassium and nitrogen and low concentration of salt.
  • A new alien species in Hungary: Limnobium laevigatum (Hydrocharitaceae)
    9-15
    Views:
    257

    The South American Spongeplant (Limnobium laevigatum (Humb. & Bonpl. ex. Willd.) Heine) is a floating aquatic plant native to freshwater habitats of tropical and subtropical Central and South America. It is frequently used for ornamental purposes in ponds and aquariums, and became invasive in the recent decades around the world. It has a high reproductive potential and a high dispersal capacity as well. It can form massive floating mats causing light limitation and creating anoxic conditions in the underlying water column, which strongly reduces native animal and plant biomass and diversity. It can also hamper navigation and water flow in rivers and canals. It has been introduced to the United States, Australia, Indonesia, Japan, Zambia and Zimbabwe. There is only one previous record in Europe (Belgium). This article is about the first record of Limnobium laevigatum in Hungary. I found two localities situated near Tata-Naszály and Dunaalmás (North-western part of Hungary). Both localities are fed by hot-water springs. It forms a small but dense population in Dunaalmás located near to the hot spring. The population in Tata-Naszály can be found in a 1.3 km long section of a stream, where it formed a sparse population in 2018.

  • Data to the flora of Hungary I.
    254-259
    Views:
    79

    The paper gives new floristic data from several regions of Hungary. Some of them are recent evidences of old data (e.g. Herniaria incana in Pomáz), others are new occurrences of recently expanding alien species (e.g. Eleusine indica, Geranium purpureum, Typha laxmannii), casual species (e.g. Lobularia maritima, Nepeta racemosa) or legally protected taxa (e.g. Agrostemma githago, Ornithogalum brevistylum, Ranunculus illyricus, Sonchus palustris).

  • Floristic data from Lengyel–Hőgyész Natura 2000 site (SW Hungary)
    31–38
    Views:
    164

    This paper presents occurrence data of 169 taxa from the Lengyel–Hőgyész Natura 2000 site (HUDD20026) which is on the southern part of Tolna Hills, SW Hungary. Data were collected during three years (2015, 2016, 2017), on different sites annually. The presented data contribute to the distribution dataset of Atlas florae Hungariae. Regionally rare or sporadic species (e.g. Asplenium scolopendrium, Asplenium trichomanes, Polystichum setiferum, Doronicum hungaricum, Paris quadrifolia, Scilla vindobonensis) are enumerated. Presence of Epipactis leptochila subsp. neglecta and Platanthera chlorantha were not registered from this site before. Subspecies of Carex divulsa are also studied on this site – distribution of these infraspecific taxa have not been mapped recently in Hungary.

  • The purple viper's bugloss (Echium plantagineum), a new adventive species for Hungary
    199-206
    Views:
    886

    Following global trends, the number of newly established alien plant species is increasing in Hungary. Not only professional scientists but also citizens could contribute to the discovery and documentation of new occurrences. One of the first records of the purple viper’s bugloss (Echium plantagineum L.) in Hungary originate from an online group dealing with wild plant species identification. This species is native in Western Mediterranean countries, and is introduced to other continents. In Australia, it is a well-spread, largely problematic invasive species, and it might show similar tendencies in Hungary as well. In order to acquaint the wider public with this species, we present its morphology, phenology and other characteristics and provide a possible inlay in the Hungarian identification key.

  • Epipactis pseudopurpurata Mered’a, a new orchid species for the Hungarian flora
    27–32
    Views:
    111

    A small population of Epipactis pseudopurpurata new to the flora of Hungary was found in the Buda Mts, near Telki village (Pest county). The main diagnostic features of this species are discussed, and a comparison with the allied species, E. purpurata is provided.

  • Distribution and identification key of Rubus L. ser. Micantes in Hungary
    56-68
    Views:
    82

    Based on a substantial field-work and herbarium revision carried out between 2007 and 2014, the authors present the taxonomical evaluation of recently known representatives of Rubus L. ser. Micantes Sudre in Hungary, providing the first grid-based distribution maps and an identification key of four species of the section (Rubus balatonicus Borbás, R. clusii Borbás, R. styriacus Halácsy, R. tabanimontanus Figert) for Hungary.

  • The occurrence of Viola collina Besser in the Bükk and Uppony Mts. (N Hungary)
    254-258
    Views:
    157

    Herbarium revisions and recent field observations revealed the occurrence of Viola collina Besser in the Bükk and Uppony Mts. (northern Hungary). This species has not been reported from the North Hungarian Mountains thus far. Altogether 94 localities of V. collina have been recorded in that region between 2007 and 2014. A map of the species’ local distribution is provided. Local habitat preferences of V. collina are briefly discussed.

  • Historical occurrence of Spergularia echinosperma, a rare Nanocyperion species in Hungary
    3–9
    Views:
    297

    Three species of the genus Spergularia have been known from Hungary. The most widespread one, Spergularia rubra (L.) J. Presl et C. Presl occurs on young sandy fallow fields and other disturbed habitats.  The other species, S. marina (L.) Bess. and S. media (L.) C. Presl, occur sporadically in saline grasslands, and both have been recently spreading along salted roads. During a revision of the genus Spergularia in Hungarian herbaria, specimens of the species S. echinosperma (Čelak.) Asch. et Graebn. were found. It is a rare species previously known to occur only in Czechia and Germany and to be extinct in Austria and Slovakia (a few occurrences along the Morava and Danube rivers between 1909 and 1953). However, it was recently discovered in Western Siberia and Kazakhstan. It is a typical mudflat plant that occurs on exposed sandy shores of water bodies. Two herbarium sheets collected by Gyula Tauscher from near the village of Ercsi (riverbank of the Danube) in 1903 were found in the Carpato-Pannonicum collection of the Hungarian Natural History Museum. In addition, a new specimen was found, collected in Slovakia (Danube near Šamorín, collected by Mihály Résely in 1868) at the vicinity of the Hungarian border. The species was not confirmed during a field inspection of the target localities in Hungary; therefore, we have to consider it extinct in the country as well as in the whole Pannonian Basin. Recently, S. echinosperma was split into two subspecies – subsp. echinosperma and subsp. albensis. The subsp. echinosperma is probably restricted to fishponds in the SW part of Czechia and subsp. albensis occurs mainly in the floodplains of the river Elbe in Germany. The extinct populations of S. echinosperma in the Pannonian Basin belonged to the subsp. albensis. This taxon is morphologically close to S. rubra but can be reliably distinguished by densely verrucose seeds and shorter than wide stipules.

  • Some floristic data from surroundings of Kötcse II. (West Outer Somogy, South Transdanubia, Hungary)
    114-121
    Views:
    110

    Floristic data of 73 species from NE part of West Outer Somogy are provided in the paper. Most of them were collected in 2016. The paper contains distribution data of species, which have not been detected in Outer Somogy so far (e.g. Peucedanum arenarium), and new occurrences of rare or sporadic taxa (e.g. Menyanthes trifoliataVeronica scardica), as well as subspontaneous occurrences of cultivated species (e.g. Narcissus poëticusHelleborus purpurascens) are provided.

  • Cyperus odoratus in Hungary
    157–164
    Views:
    323

    Eleven species of the genus Cyperus have been reported from Hungary so far, of which 3 are certainly aliens. A small population of C. odoratus, new for the flora of Hungary, was found along the River Danube near Foktő settlement (Bács-Kiskun County, S Hungary) in late autumn of 2019. On this occasion blooming specimens were observed in the floodplain ruderal vegetation on the bank of a fork of the Danube. This species is widespread in the tropical and subtropical regions of the World, and was first introduced to Europe in the early 1950s.  More recently it has been spreading along certain rivers of Italy and Spain. It was discovered along the river Danube in Romania in the early 1990s, and later at the upper parts of the river in Bulgaria, Croatia and Serbia. Its introduction into Hungary is connected with the Danube, too. Based on foreign experience, it is probable that C. odoratus will be spreading along the Hungarian Danube system, hopefully, not as an invasive species.

  • Contributions to the lichen flora of the Mecsek Mts (South Hungary)
    16–26
    Views:
    95

    A checklist of 115 lichen taxa was compiled from the Mecsek Mts, based on floristical records of 221 old or recently collected herbarium specimens (incl. 37 duplicates) deposited in the JPU herbarium (Pécs, Hungary). The occurrence of 22 taxa are new for the Mecsek Mts: Arthopyrenia cinereopruinosaBaeomyces rufusCandelariella reflexaCatapyrenium rufescensCladonia macilentaDimerella pinetiHypogymnia tubulosaOchrolechia arboreaPleurosticta acetabulumPseudevernia furfuraceaPsilolechia lucidaPycnothelia papillariaRinodina sophodesTomasellia arthonioidesToninia candidaTrapelia involutaVaricellaria hemisphaericaVerrucaria marmoreaV. nigrescensV. parmigeraXanthoparmelia conspersa, X. verruculifera. One legally protected lichen species (Peltigera leucophlebia) is also represented by two specimens.

  • The occurences of two ‘old-new’ alien grass species, Rostraria cristata (L.) Tzvelev and Cynosurus echinatus L. in Pécs
    39-42
    Views:
    86

    Some small populations of Rostraria cristata and Cynosurus echinatus, two ‘old-new’ alien grass species in Hungary, were recently discovered in the city of Pécs (South Hungary). A morphological description of these species as well as a survey of their former and current localities are presented. 

  • Floristic data from the Great Plain of Hungary (Alföld)
    304–316
    Views:
    226

    In this study we report relevant occurrence data of 49 species and hybrids and their habitats from the Great Plain registered in the period between 2011 and 2017. Due to importance for nature conservation, we present additional data of further 13 species. Most of the data are originated from riverine oak-elm-ash forests of South East Hungary (from the region of Gyula, Békéscsaba, Doboz settlements). The most important result of our study is the rediscovery of Gagea minima and Dictamnus albus. Besides, we report new occurrence data for the Great Plain (Alföld), as well as for the flora of the microregions in questions (e.g. Draba muralisNicandra physalodesCystopteris fragilis) and also present the data of the regionally endangered species (e.g. Inula heleniumOphioglossum vulgatum). Some rare weed taxa (e.g. Calepina irregularisCardamine impatiens), spreading adventive taxa (e.g. Chorispora tenellaEuphorbia maculataPhytolacca americanaPh. esculentaSicyos angulata) and in surveys rather underrepresented species (e.g. Loranthus europaeus, Arabis hirsutaRumex confertus) are reported too.

  • A new montane bryophyte species from the Mátra Mts, Hungary: Hymenoloma crispulum
    11-18
    Views:
    233

    Due to the geomorphological situation and climatic conditions of Hungary, true montane climate is restricted to very few and small areas of higher elevation, with the highest peak (Mt Kékes, 1014 m) in the Mátra Mountains (North Central Hungary). During a bryofloristical survey in this re­gion, a few cushions of a Boreo-arctic montane bryophyte species, Hymenoloma crispulum (Hedw.) Ochyra (syn.: Dicranoweisia crispula (Hedw.) Milde), was discovered at the steep northern slope of Mt. Kékes in an andesite boulder scree with cool and humid microclimate. H. crispulum is new to the Hun­garian bryoflora and fits in well with other co-occurring vascular and cryptogamic taxa of northern-montane affinities. The population is very small and vulnerable, therefore it should be rated 'critically endangered' (CR) in an updated national red list.