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Online distribution atlas of the Hungarian vascular flora (Atlas Florae Hungariae): Homepage structure and prospects of database building
238–252Views:925In December 2018 the online Distribution atlas of vascular plants of Hungary (Atlas Florae Hungariae) was published. The database was built from more than 1 million data records. It mainly contains data from the Hungarian Flora Mapping Programme, but herbarium and literature data were also processed. Maps of species richness illustrate the taxonomic diversity of biogeographical areas and shows the differences of survey quality. After the Flora Mapping Programme ended, between 2016 and 2019, additional data from scientific articles were processed. 25% of new data originated from the article series “Contributions to the Atlas Florae Hungariae” in the Kitaibelia journal, which significantly broadened the distribution of some species. Records of Mesophile forest species (e.g. Allium ursinum, Dryopteris dilatata) increased, as well as records of weed species (e.g. Euphorbia maculata, Senecio vernalis) which can be attributed to their natural spreading. The most important future tasks are to create a reviewed list of taxa and extend data records with an attribute table containing all additional information. Until now the database website is presented only in Hungarian (http://floraatlasz.uni-sopron.hu). Registration is available for anybody, after sending a request via e-mail (atlas.florae.hungariae@uni-sopron.hu). There are different options (excel file by e-mail or online web form) on how to upload floristic data for registered users. Data is only accepted if required information (name of taxa, date, data publisher, settlement, CEU code) is filled out. Each record of uploaded data is revised by the website administrator and taxa specialists. The English version of the webpage will be available in the near future.
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Data on the flora of the burial mounds of the Great Hungarian Plain
31–44Views:357In this paper, we present plant occurrence data from ancient burial mounds, located in the Danube–Tisza Interfluve (Praematricum and Colocense) and in the Trans-Tisza region (Crisicum), Hungary. We report data on 45 vascular plant species and one fungus (Morchella steppicola) from a total of 109 mounds. Of the species presented, 13 taxa are protected in Hungary, some of which are not rare in the study region (e.g. Aster sedifolius, Astragalus asper, Linaria biebersteinii, and Ornithogalum brevistylum). We present new occurrence records of some spreading annual species (e.g. Aegilops cylindrica, Crepis pulchra, and Trifolium diffusum), as well as data on some species that have not been previously reported from mounds (e.g. Agrostemma githago, Chaerophyllum bulbosum, and Dryopteris filix-mas), and we also publish the occurrences of an alien species (Lolium multiflorum) on burial mounds. In terms of plant biogeography, the occurrence of Orobanche reticulata in Kengyel and Oxytropis pilosa in Tömörkény is considered an especially important data.
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Floristic records from Northern Hungary III. Zemplén Mts
109–172Views:753We report new floristic data collected from the Zemplén Mountains between 2004–2022. Altogether, 1215 occurrences of 133 species were recorded from 33 Central European Flora Mapping quadrats. We present occurrences of so far not reported or rare species, e.g., Carex appropinquata, C. cespitosa, C. disticha, Centaurea stenolepis, Scabiosa columbaria, Tephroseris aurantiaca. We put special emphasis on occurrence data of geobotanical importance, such as Anthriscus nitidus, Echium maculatum, Lunaria rediviva, Nepeta pannonica, Oreopteris limbosperma, Petasites albus, Primula elatior, Thlaspi jankae, Veronica montana, as well as newly rediscovered species based on archive data: Cirsium rivulare, Parnassia palustris. We mention common species, which are lacking data from the area, e.g., Adoxa moschatellina, Allium flavum, Cardamine amara, Dipsacus fullonum, Galanthus nivalis. Moreover, we report numerous new occurrences of taxa typical to the Zemplén Mts.: Aruncus dioicus, Huperzia selago, Lycopodium annotinum, Securigera elegans etc. We complete the enumeration with additional data from the literature and herbaria.
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Contributions to the Atlas Florae Hungariae VI.
87-102Views:725The current paper is the sixth one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 389 vascular plant taxa from 117 flora mapping quadrates (CEU) are provided. New records are indicated from different regions of Hungary; however, most occurrences are located in the North Hungarian Mts and from the city of Szolnok and Western Hungary. Occurrence data of rare native taxa (including data of Equisetum fluviatile, Galium rivale, Gladiolus imbricatus, Hypericum maculatum, Persicaria bistorta, Potentilla inclinata, Ranunculus lingua, Rhinanthus rumelicus, Trifolium fragiferum subsp. bonannii, Viola pumila), rare or data-deficient alien taxa (e.g. Amaranthus blitum subsp. blitum, Cymbalaria muralis, Elymus elongatus, Euphorbia peplus, Heracleum sosnowskyi, Impatiens balfourii, Lepidium densiflorum, Rhus typhina, Telekia speciosa), as well as frequent but more or less underrepresented taxa (e.g. Carex otrubae, Chenopodium polyspermum, Crepis tectorum, Filipendula vulgaris, Petrorhagia prolifera, Poa palustris) are also enumerated. In case of the most common species we provided the CEU codes only.
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Some floristic data from surroundings of Kötcse (South Transdanubia, West Outer Somogy)
74-80Views:360There are very few published and documented floristic data from the hills of the SW region of Hungary called 'Outer Somogy'. This paper present the occurrence data of 31 vascular plant species from the western part of this region. Among them 14 taxa are legally protected and further 17 taxa are sporadically occurring. The records were collected within the administrative area of villages Karád, Kereki, Kötcse, Nagycsepely, Szólád (Nezde) and Teleki. Most data were recorded in 2014. This is the first report on the occurrence of 1) Neotinea ustulata subsp. aestivalis in South Transdanubia, and 2) Epipactis microphylla and E. tallosii in Outer Somogy. Data provided on the occurrence of Cirsium boujartii on the northwestern border of the distribution area of this insufficiently known species.
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Contributions to the Atlas Florae Hungariae III.
122-146Views:2024The current paper is the third one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 558 vascular plant taxa from 186 flora mapping quadrates (CEU) are presented. New records are indicated from different regions of Hungary; however, most occurrences are located in the North Hungarian Mts. Occurrence data of rare native taxa (e.g. Marsilea quadrifolia, Salicornia prostrata, Stellaria palustris, Potentilla patula, Althaea cannabina, Lythrum tribracteatum, Ajuga laxmannii, Plantago schwarzenbergiana, Alisma gramineum, Gagea bohemica, G. szovitsii), rare or data-deficient alien taxa (e.g. Chorispora tenella, Thladiantha dubia, Senecio vernalis) as well as frequent but more or less underrepresented taxa (e.g. Scleranthus annuus, Vicia hirsuta, V. lathyroides) are also enumerated. In case of the most common species we provided the CEU codes only.
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Contributions to the Atlas Florae Hungariae II.
227-252Views:945The present article is the second part of the series aiming to contribute with new data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 612 vascular plant taxa from 283 flora mapping quadrates (CEU) are presented. New records are distributed over the entire Hungary, however, most are localised in the North Hungarian Mts. Occurrence data of rare native taxa (e.g. Antennaria dioica, Bupleurum pachnospermum, Carex rostrata, Ceratocephala testiculata, Cicuta virosa, Cirsium boujartii, Epipactis moravica, Lycopsis arvensis, Parnassia palustris, Pyrus nivalis, Rumex kerneri, Rumex pulcher, Silene nemoralis, Stellaria alsine, Teucrium botrys, Vicia lutea), rare or data-deficient alien taxa (e.g. Nonea lutea, Panicum dichotomiflorum, Silybum marianum) as well as frequent but more or less underrepresented taxa (e.g. Eragrostis minor, Heliotropium europaeum, Saxifraga tridactylites) are also enumerated, since our intention was to fill the gaps in the Atlas. Remarcable contribution on the occupied territory of Ranunculus illyricus and Spergula pentandra in the Nyírség region (E Hungary), and on the Hungarian distribution of expanding alien weed Senecio vernalis are also presented.
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Chorological notes on the vascular flora of Hungary
85-119Views:685The paper presents 1593 current chorological data of 584 taxa (species, subspecies, varietas, hybrids) from Hungary, collected during the last few years. All of these connected also to a total of 184 CEU quadrats. Among the species characteristic to the natural habitats, we highlighted the records of several species protected in Hungary (Adoxa moschatelina, Althaea cannabina, Cardamine parviflora, Centaurium pulchellum, Gagea pusilla, Orobanche reticulata, Potentilla patula, Rosa zagrabiensis, Spergula pentandra, Trifolium diffusum, Valerianella dentata, V. rimosa, Veronica officinalis, Viola collina and V. pumila). In the Northern part of Hungary, we observed the establishment of Cerastium dubium and Ranunculus pedatus at several places along roadsides, which species have been known from the surrounding saline areas for a long time, but their appearance at roadsides is relatively new phenomenon. In addition, Aegilops cylindrica, Crypsis schoenoides, and Fumaria vaillantii subsp. vaillantii were found in one location – each species on roadsides. Along the railway tracks, we found disturbance-tolerant pioneers of rocky grasslands, open sandy and loess grasslands, e.g. many population of the native Androsace elongata, Anthemis ruthenica, Centaurea arenaria, Draba nemorosa, Erodium ciconium, Medicago monspeliaca, Microrrhinum minus, Saxifraga tridactylites and Vulpia myuros were found, as well as numerous ruderalia and the archaeophyt such as Aegilops cylindrica. In addition to the more common species from the urban flora, we present some new data on the occurrence of the rarer Amaranthus blitum subsp. emarginatus, Chenopodium glaucum, Ch. murale, Ch. opulifolium, Ch. vulvaria and Euphorbia peplus. We have also listed some of the rarer field weeds and species that can establish in younger and older old-fields. Bromus secalinus has been found in several places in the northeastern part of Hungary. We also observed Calepina irregularis, Centaurea solstitialis, Centaurium pulchellum, Erysimum cheiranthoides, Filago lutescens, Fumaria officinalis subsp. officinalis, Geranium dissectum, Misopates orontium, Myagrum perfoliatum, Reseda phyteuma, Spergularia salina and Veronica anagalloides populations. Erodium hoefftianum is a member of the natural flora, but is now found only in old-fields. We found some pasture weeds in regions where they are not common (e.g. Bupleurum tenuissimum, Cerastium dubium and Trifolium striatum). The study also presents some species that are data deficient for taxonomic reasons, but not very rare ones (or we don’t know that). This group includes Arabis sagittata, Bolboschoenus glaucus, Bromus commutatus, B. racemosus, B. ramosus, Fumaria vaillantii subsp. schrammii, Ononis spinosiformis subsp. semihircina, Potentilla collina and Viola suavis “white morfotype”. We have drawn attention to the effect of ecological-botanical works on flora-influencing. We found Anthemis ruthenica (species of sandy grasslands) in a mountain meadow, which is probably the result of seeds accidentally introduced by researchers. Finally, the paper provides data on the current distribution of hybrid taxa. A new individual of Betula ×rhombifolia was established in the Mátra Mts. We found some Euphorbia individuals that appeared to be a transitional form between E. esila and E. virgata (E. ×intercedens; Eu. esula subsp. saratoi). We also present some Verbascum and Viola hybrids. Hybrids were determined on a macromorphological basis.
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Contributions to the flora of the Szuha watershed and adjacent territories (NE Hungary)
27-67Views:495This paper presents new floristic data of 415 taxa from the Putnok Hills and Sajó Valley microregions (Northern Hungary). Records were collected between 1991 and 2021. In the first part new occurrences of legally protected (e.g., Cypripedium calceolus, Iris aphylla subsp. hungarica, Lycopodium clavatum, Thalictrum aquilegiifolium, Pyrola rotundifolia, Scorzonera purpurea, Stipa pulcherrima) and not protected but locally rare or data-deficient species (e.g., Myosurus minimus, Thalictrum simplex, Sisymbrium loeselii, Camelina microcarpa, Potentilla collina, Trifolium striatum, Geranium pratense, G. dissectum, G. divaricatum, Viola pumila, Epilobium roseum, Myriophyllum spicatum, Pimpinella major, Berula erecta, Trinia ramosissima, Centaurium pulchellum, Nepeta cataria, Limosella aquatica, Veronica scutellata, Orobanche arenaria, Utricularia vulgaris, Galium elongatum, Cephalaria transsylvanica, Dipsacus fullonum, Xanthium saccharatum, Alisma lanceolatum, Elodea canadensis, Potamogeton pectinatus, P. berchtoldii, P. lucens, P. nodosus, Zannichellia palustris, Sisyrinchium bermudiana, Melica picta, Elymus elongatus, Leersia oryzoides, Typha laxmannii, Eriochloa villosa) are presented. This chapter contains data on 246 species from which 8 are new for the flora of the Tornense flora region and 47 for its subregion, the Putnok Hills. Further contributions to the distribution data of Atlas Florae Hungariae are presented in Electronic Appendix. This section contains raw occurrence data of 169 species.
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Data on the native vascular plant species of the Pannonian Region II. (6–22)
233–252Views:956In the second part of this article series, which provides data contributing to the knowledge of native plant species of the Pannonian Ecoregion, we address rare species – mostly legally protected in Hungary – belonging to the families Cephaloziaceae, Athyriaceae, Dryopteridaceae, Cystopteridaceae, Thelypteridaceae, Brassicaceae, Rosaceae, Elatinaceae, Apiaceae, Orobanchaceae, Liliaceae, and Orchidaceae. We report the current occurrence of Nowellia curvifolia in the Sub-Pannonian region of Slovenia. We present an exceptionally species-rich habitat for ferns (Athyrium filix-femina, Dryopteris carthusiana, D. dilatata, D. filix-mas, Gymnocarpium dryopteris, Polystichum aculeatum) from the ‘Borsodi ártér’ microregion. We provide data on the stands of Thelypteris palustris observed along the main irrigation canals of ‘Tiszántúl’ region. We publish new occurrence records contributing to the Hungarian distribution of Aethionema saxatile, Crataegus nigra, Elatine hungarica, Sium sisarum, Orobanche hederae, Himantoglossum adriaticum, Epipactis placentina, E. exilis, E. pseudopurpurata, and E. microphylla, as well as to the appearance of Helosciadium repens in urbanized environment. Gagea spathacea and G. bohemica are newly recorded for the Zemplén Mts; Epipactis nordeniorum and Epipogium aphyllum are new for the flora of the Mátra Mts; and Himantoglossum calcaratum is newly recorded for Hungary. We also present spontaneous establishment of several orchid species in the courtyard of a secondary school in Veszprém. Additionally, we report on the pollinating insects observed on the flowers of Sternbergia colchiciflora. Two previously misreported records are also corrected.
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Contributions to the Atlas Florae Hungariae XI.: Data from the Tokaj–Zemplén Mts and its surroundings
239–241Views:376This paper is the 11th in the series aiming to contribute data to the distribution maps published recently in Atlas Florae Hungariae. Our records were collected from the Tokaj-Zemplén Mts and the Bodrogköz region. The dataset includes 325 occurrence data of 75 vascular plant taxa from 53 flora mapping unit. Most of the represented taxa are legally protected, e.g. Aconitum variegatum, Alchemilla subcrenata, Aster amellus, Cardamine glanduligera, Centaurea triumfettii, Cephalanthera damasonium, Dianthus deltoides, Epipactis pontica, Gentianopsis ciliata, Orchis morio, Orthilia secunda, Petasites albus, Platanthera bifolia, Polygala major, Prunella grandiflora, Pulsatilla grandis, Rosa pendulina, Salix aurita, Scilla kladnii and Stipa dasyphylla.
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Data to the flora of Sopron Hills and its foreground
239-242Views:473The present study reports additional records to the flora of Sopron Hills and Sopron Basin. The records were collected during our field surveys between 2010 and 2014 and contain occurrence data of 19 vascular plant taxa. The genus Orobanche is represented by 5 species. Two taxa are new for the flora of Sopron Hills (Amaranthus blitoides, Orobanche teucrii), others are interesting because of their old or uncertain indication (e.g. Dianthus superbus, Myosotis sparsiflora, Orobanche lutea, Orobanche purpurea).
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Floristic records from Northern Hungary II. North Cserehát and Upper Bódva valley
31–48Views:444This paper presents new floristic data of 123 taxa from the regions of North Cserehát and Upper Bódva valley (Northern Hungary). Records were collected between 1995 and 2020. New occurrences of legally protected (e.g., Dactylorhiza incarnata, Eriophorum angustifolium, Lycopodium clavatum) and not protected but locally rare or data-deficient species (e.g., Artemisia annua, Carex remota, Cirsium pannonicum, Cyperus fuscus, Genista pilosa, Melampyrum cristatum Ranunculus flammula, Scorzonera purpurea, Silene noctiflora) are presented.
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New species and changes in the bryophyte flora of the Mecsek valleys (Hungary)
157–176Views:328The paper discusses the bryophyte flora of 77 sites in the Mecsek Mountains, mainly in deeper valleys. It includes species that are new to the area’s flora and/or endangered in Hungary, as well as mosses that have become more widespread or locally abundant compared to data from 1968 and 1983. The names of the locations, their geographical centers (centroids), and the grid unit IDs of the Hungarian Flora Mapping Programme (HFMP), where the sites are located, are listed and referenced according to their number in the list. Some of the species are national rarities, such as Scleropodium touretii, newly recorded in the Mecsek in 2023. Its second known occurrence in Hungary is reported in the Enumeration. We detail the occurrences of several species new to South Transdanubia, including Dicranum spurium, Diplophyllum albicans, Hypnum jutlandicum, Isothecium myosuroides, Kindbergia praelonga, Rhynchostegiella curviseta, Riccardia palmata, and Sciuro-hypnum plumosum. Additionally, species newly recorded from the Mecsek Mountains include Fontinalis antipyretica, Leptodon smithii, Plagiomnium ellipticum, Pseudocampylium radicale, and Tortula brevissima. For two taxa, we present data of importance not only for the Mecsek, but also for the North-Hungarian Mts: Riccardia palmata, new for the Tornai-karszt, and Dicranum spurium, for which we confirm a very old record in the Zemplén Mts. We also note the widespread, sometimes massive occurrence of some aquatic and subaquatic mosses, e.g. Rhynchostegiella teneriffae, Hygrohypnum luridum, which have been recorded only once in the mountains in the last 50 years. The taxonomically uncertain Hygroamblystegium fluviatile with similar habitat requirements can now be found widely throughout the mountains. New populations of Palustriella commutata, considered endangered and declining in the country, have been found in the Mecsek. The expansion of (sub-)aquatic species is particularly surprising in the light of recent hydroecological studies indicating that the streams in the Mecsek are drying up. The situation is similar regarding Rhynchostegium rotundifolium, previously thought to be a Tilio-Acerion (slope and ravine forest) species, which is legally protected in Hungary and only had one record from the Mecsek in the last century. This forest species is now found in peri-urban, sub-urban, slightly to moderately disturbed and secondary habitats throughout the mountains. Similar to its preference in Germany, outside of semi-natural habitats it occurs mainly on built structures in forest edges, ruins, abandoned mine sites, often in stands mixed with black locust and contaminated with litter. Microeurhynchium pumilum, which prefers sub-Mediterranean and sub-Atlantic climate, shady and humid habitats, has become one of the most widespread and abundant forest bryophytes in the Mecsek, in contrast to its few records 60 years ago. Half (50%) of the new mosses in the flora of the Mecsek can be connected to the cooler boreal biome, while 21% is associated with the milder Mediterranean-Atlantic biome. The average of MAT (mean annual daily mean air temperatures averaged over one year) and MAP (accumulated precipitation amount over one year) of the 14 new species according to BET (Bryophytes of Europe Traits) indicates areas with much lower temperatures (7.7°C) than in Hungary, but with precipitation conditions (749 mm) similar to those in the Mecsek. The vast majority of the new species are extremely rare in Hungary, and are inconsistent with current climate warming trends. In contrast to the mosses with striking increase in records, the detection of new species is likely due to chance, in addition to systematic research at finer spatial scales, rather than to local expansion of the species. However, in the case of expanding and abundant species, it is still an open question whether their dynamics compared to 40-60 year old references are apparent or real: the result of sampling bias in previous surveys or a consequence of climate change. In any case, the abundance of the above mentioned bryophytes in the Mecsek is not consistent with their current Hungarian Red List threat classification, and this cannot be attributed to changes in definitions since 2010. The detected distribution and abundance, as well as the increase in the number of new species in the country within a decade, highlights the need for an updated Red List of Hungarian bryophytes.
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Contributions to the Atlas Florae Hungariae VII.
218-237Views:408This paper lists supplementary data to the recently published Atlas Florae Hungariae, mostly from various parts of Transdanubia and the Great Plain. The list includes 73 plant species with at least one new occurrence previously not shown on their grid-based range map, as well as data confirming old records or correcting erroneous distribution data. The actual habitats where the plants occur are characterized by their corresponding ÁNÉR codes.
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Data on alien vascular plant species in Hungary II. (7–12)
253–264Views:608In the second part of our series providing data on alien plant species in Hungary, we address taxa belonging to the families Cabombaceae, Brassicaceae, Rubiaceae, Buxaceae, Violaceae and Compositae. We present new records of the distribution of Cabomba caroliniana on the Körös Plain: along the Élővíz Canal between the towns of Békés and Békéscsaba the species was found at 17 sites, covering a total area of more than 630 m² in slightly alkaline, low-conductivity water that remains unfrozen in winter. In Szentendre, we document a transient escape of Isatis tinctoria and several populations of Rubia tinctorum, some of which can be traced back to former cultivation. We report the first extra-garden, woodland occurrence of Pachysandra terminalis in Hungary and relate it to the dumping of green waste. We clarify the status of Viola prionantha in the Hungarian flora and list numerous occurrences mainly associated with urban pavements, parks and cemeteries. We detected a new occurrence of the Atlantic halophyte Cochlearia danica on the Great Hungarian Plain, near Karcag, along the Main Road 4. Finally, we describe the first Hungarian record of Symphyotrichum squamatum from pioneer silt surfaces along a canal near Szeged, highlighting its potential invasive role.
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Fern (Pteridopsida) flora of South Nyírség (East Hungary)
162–182Views:594The study presents a number of new data on the fern flora of South Nyírség, and reviews all available literature and herbarium fern records from the area. A total of 19 Pteridopsida species were detected in the region. The hart's tongue fern (Asplenium scolopendrium) and the soft shield fern (Polystichum setiferum) were hitherto unknown in the whole Nyírség region (flora district “Nyírségense”). The broad buckler-fern (Dryopteris dilatata), scaly male fern (Dryopteris affinis) and floating fern (Salvinia natans) are new records to the area of South Nyírség. Dryopteris dilatata has been found in a surprisingly great number of localities. Some morphological characters facilitating identification of this species are also discussed. The local pine plantations, fens, carrs and humid deciduous forests are important habitats for ferns. In some localities the co-occurrence of 5-6 fern species were observed. The increasing aridity in the last decade is posing a serious threat to ferns in the area.
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The historical occurrence of Digitaria ciliaris in Hungary – results of a herbarium survey
53–62Views:473Revision of genus Digitaria in BP Herbarium Carpato-Pannonicum collection helped to discover the historical occurrence of Digitaria ciliaris (Retz.) Koeler from Győr, in the area of an old cotton and wool processing and textile factory. However, it has been proved that the previously published Hungarian records of the species are all wrong, as the Digitaria-keys of Hungarian herbals misinterpreted the taxon, mainly as a synonym of D. sanguinalis subsp. pectiniformis. The herbarium survey confirmed that D. sanguinalis is the most common taxon in Hungary. D. sanguinalis subsp. pectiniformis is also widespread, but probably somewhat under-mapped. Probably not native to Hungary, most probably an archeophyte taxon. Specimens have been collected from disturbed habitats associated to railways and from xerothermic vineyard areas for over a hundred years. D. ischaemum is certainly native to the region, with a recognisable natural distribution pattern, and is found mainly in the sandy and gravelly soils of the lowland and hilly areas of the country. This species has recently become more common in urban areas across the country with the recent spread of sand-lined pavements. D. ciliaris of tropical/subtropical origin is a casual alien species that was formerly present in the Hungarian flora.
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Floristic records from Northern Hungary IV. Helleborus dumetorum from the Cserhát Hills and other occurrences
209–210Views:415In this publication, the author provides occurrences of geobotanical importance from the North Hungarian Mountains collected in the spring of 2023. First occurrence for the following species was recorded: Helleborus dumetorum from the Cserhát Hills, Lunaria annua from the Mátra Mts. and Dryopteris affinis from the Bükk Mts. Ranunculus aquatilis was found again in the Bükk Mts., where it had been collected more than 100 years ago, and Rubus saxatilis was found for the fourth time in the Zemplén Mts.