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75-78.
Vol 15No 42009

Still a large amount of pesticides and spary applications are used in environmentally friendly fruit production systems; therefore, the aim of our study was first to test the in vitro effeicacy of some fungicides against a key apple disease (apple scab), and secondly to evaulate the effectiveness of reduced spray programmes against apple s...cab, powdery mildew and coding moth in integrated and organic apple orchards. In vitro efficacy of 7 fungicides (Champion 50 WP, Kocide 2000, Nordox 75 WG, Olajos rézkén, Kumulus S, Rézkén, Rézoxiklorid) and another 6 fungicides (Score 25 EC, Efuzin 500 SC, Systane, Folicur Solo, Zato Plusz, Rovral) approved in organic and integrated production systems, respectively, were tested against apple scab. Altogether four spray programmes were performed i) standard integrated: sprays followed by forecasting systems during the season, ii) reduced integrated: sprays followed by forecasting systems but only 75% of the spray numbers used during the season-long spray programme, iii) standard oragnic: sprays applied every 7–14 days during the season and iv) reduced organic: 60% of the spray numbers used during the season-long spray programme. In vitro results showed that fungicides (with active ingredients of copper and sulphur) applied in organic production showed relatively high percent growth capacity of the apple scab fungus. Rézkén showed the highest and Kumilus S the lowest efficacy against apple scab. Fungicides applied in integrated production showed relatively low percent growth capacity of the apple scab fungus. Score 25 EC showed the highest and Rovral the lowest efficacy against apple scab. Field study showed that reduced spray programmes did not increase significantly scab incidence in the integrated field. However, scab incidence increased significanly (above 30%) in the reduced spray programme for the organic orchard.Mildew incidence was low (below 5%) in both integrated and organic spray programmes. Mildew incidence on both shoots and fruits increased significanly in the reduced spray programme for the organic orchard. Incidence of codling moth damage was affected the most by standard vs. reduced spray programmes. Though incidence remained below 10% in the integrated plots, the incidence was significantly higher in the reduced spray programme compared to the standard programme. Similar results were obtained for organic spary programmes, but the incidence was 10 times higher and the differences among the two programmes were larger.,

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85
85
19-23.
Vol 16No 32010

Fruits of different sour cherry varieties cultivated, in 2008 and 2009, under organic farming and integrated cultivation conditions were analysed for their quality attributes, antioxidant activity and subjected to sensory evaluation.. Average size, weight, soluble solids, titratable acidity, total polyphenols, free radical scavenging capacity e...xpressed as Trolox equivalent (TEAC), copper and zinc were determined in freshly harvested fruits. The obtained results indicated that, the principal component analysis can separate and distinguish the seasons of fruit production. The farming system seemed to have slight effect on quality the fruit as compared to varietal factors (genotypes). However, the total polyphenol content was uniformly less in 2009. Total polyphenols and free radical scavenging activity were significantly higher in Bosnian type sour cherries, and outstanding in Amarelle type cultivar ‘Pipacs’. There was no statistically significant difference between the sensory properties of cultivars tested by panels, except the case of ‘Pipacs’. The organolaptic investigation showed marked preference to the fruits of Eva and Petri cultivars.

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50-53.
Vol 16No 12010

The aim of this study was first to test the in vitro effeicacy of some fungicides against brown rot of sour cherry, and secondly to evaulate the effectiveness of reduced spray programmes against brown rot in integrated and organic sour cherry orchards. In vitro efficacy of 7 fungicides (Champion 50 WP, Kocide 2000, Nordox 75 WG, Olajos rézkén..., Kumulus S, Rézkén, Rézoxiklorid) and another 6 fungicides (Score 25 EC, Efuzin 500 SC, Systane, Folicur Solo, Zato Plusz, Rovral) approved in organic and integrated production systems, respectively, were tested against brown rot of sour cherry. Altogether four spray programmes were performed i) standard integrated: sprays followed by forecasting systems during the season, ii) reduced integrated: sprays followed by forecasting systems but only 75% of the spray numbers used during the season-long spray programme, iii) standard oragnic: sprays applied every 7–14 days during the season and iv) reduced organic: 60% of the spray numbers used during the season-long spray programme. In vitro results showed that fungicides (with active ingredients of copper and sulphur) applied in organic production showed relatively high percent growth capacity of Monilinia fungus. Rézkén showed the highest and Kumilus S the lowest efficacy against brown rot. Fungicides applied in integrated production showed relatively low percent growth capacity of Monilinia fungus. Score 25 EC showed the highest and Rovral the lowest efficacy against brown rot. Field study showed that reduced spray programmes did not increase significantly brown rot incidence in the integrated field. However, brown rot incidence increased significanly (above 30%) in the reduced spray programme for the organic orchard.

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69
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15-18.
Vol 22No 1-22016

Success of apple production is highly influenced by the applied production system and the planted cultivar. In this paper growing characteristics of 39 apple cultivars were studied in integrated and organic production systems. These kind of parameters are less studied in the cultivar and training system examinations, although they have huge eff...ect on the training and maintaining of canopy, on the pruning necessity, ultimately on the production costs. According to our results the thickness of the central axis of apple trees showed significant differences between the integrated and the organic systems. Axis of the trees with lower trunk thickness tapers more slightly in the integrated production system, than in the case of the trees with thicker trunk in the organic system. Thicker axis is not accompanied by thicker trunk, namely the thickness of the central leader starts to decrease stronger in the organic production system, compared to the integrated one.

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61-63.
Vol 15No 42009

The aim our study was to establish whether significant differences in nutrients uptake and quality of fruit exist
between organic and integrated grown apples. The study was performed at the orchard Fruit Research Station, University of Debrecen, at Debrecen-Pallag during 2002–2004. Diameter (mm), weight (g), firmness (N/cm2), dry matter (%...) and Vitamin C in fruit were determined. Diameter data showed that the size of apples strongly depended on cultivars and organic apples were larger than integrated ones. Moreover, the effect of year and species strongly affected the apple diameter, size and weight. Measured data of firmness were good agreement in data of diameter and weight. Larger fruits have lower values of firmness due to the structure of fruit flesh. Dry matter content of apples varied between 14.66 and 18.07 in integrated, and 12.87 and 17.29 in organic apples according to cultivars. Values were affected by years and cultivars. From results it was evident that the dry matter content of apples affected by production system. It was lower in organic samples than integrated ones.Vitamin C content of apples was stronger affected by species and years than production system.

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76
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49-51.
Vol 17No 1-22011

The aim of this study was to evaluate the effectiveness of reduced spray programmes against cherry leaf spot in integrated and organic sour cherry orchards. Altogether four spray programmes were performed i) standard integrated: sprays followed by forecasting systems during the season, ii) reduced integrated: sprays followed by forecasting syst...ems but only 75% of the spray numbers used during the season-long spray programme, iii) standard oragnic: sprays applied every 7–14 days during the season and iv) reduced organic: 60 % of the spray numbers used during the season-long spray programme. Our study showed that cherry leaf spot did not increase above 3.1, 7.4, and 8.9% in 2008, 2009, and 2010, respectively, in the integrated orchards. However, leaf spot incidence was above 20% in all years in the organic field. Reduced spray programme did not increase significantly cherry leaf spot incidence in the integrated field in either years. However, leaf spot incidence increased significanly (above 20%) in the reduced spray programme for the organic orchard.

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101-106.
Vol 9No 3-42003

The pathogen and weather components of apple scab disease progress were analysed in a three-year study, in two environmental-friendly production systems (organic and integrated) on cvs. `Idared', `Jonica' and 'Mutsu'. Linear regression analyses of transformed disease incidence and severity data and "area under the disease progress curves" (AUDP...C) were used for the analysis of the pathogen component. To evaluate the role of the weather component in apple scab epidemic, first, the weekly disease increase was determined at a certain week (n). Weekly disease increase was related to rainfall, relative humidity, Mills' wetness period, temperature and interaction between temperature and relative humidity. Five different periods were used in the analyses: i) week (n-1), ii) week n(n-1), iii) week (n-2), iv) week (n-1)(n-2) and v) week n(n-1)(n-2). In the analyses of the pathogen component, the best transformation function was the logistic one. Regression analyses showed that disease growth rates were higher for disease incidence and for the organic production system than for disease severity and for the integrated production system, respectively. Disease growth rates for leaf incidence were higher than fruit incidence on all the three cultivars. AUDPC values showed great differences in both leaf and fruit incidences among cultivars and between the two production systems. The results the of analyses of the weather component showed that the best relationships between disease increase and weather parameters were found for fruit incidence and leaf incidence in week (n-2) in the organic and integrated production systems, respectively. Results also demonstrated that in week n(n-1) temperature played a more important role in the fungus development than the water parameters (relative humidity, rainfall and leaf wetness). Consequently, infection process is significantly dependent on almost all weather parameters, but during the incubation period the most important weather parameter is the temperature. Results were compared with similar studies and biological interpretations of the analyses are discussed.

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74
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24-29.
Vol 7No 12001

In Hungary, fruit growers are increasingly interested in environmentally oriented growing methods, such as organic and integrated systems. The main factor in creating such systems is the establishment of a system of cultivar-pruning-plant protection based on production technology. Therefore, our aim was to examine the susceptibility of apple cu...ltivars to diseases and pests and the effect of pruning technique on diseases and pests in organic and integrated growing systems.

Two pathogens (Venturia inaequalis, Podosphaera leucotricha) and two pests (Leueoptera malifoliella, Pannonychus ulmi) were observed in organic and integrated systems under "strong" and "weak" pruning techniques. Our results showed that, compared to integrated production, the pathogens and pests infected the trees more in the organic system. The pruning technique affected mainly the cultivars susceptible to diseases and pests. In general, the "weak" pruning technique correlated with a higher level of damage caused by diseases and pests than the "strong" pruning technique, especially in the organic growing system. The reason that techniques should be carefully chosen is that shoots grow faster and more vigorously after employing the "strong" pruning technique. This supports a better preservation of the trees, as a result of reduced susceptibility of plant tissues to diseases and pests.

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92
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63-67.
Vol 6No 42000

In this study, the rate of scab infection and quality parameters (content of vitamin-C, dry matter, sugar, total acids, specific weight, quality index and flesh firmness) of some apple cultivars are given, moreover, the relation between those characters was examined in two environmental systems (organic and integrated) in a rainy season. In the... organic fruit production, scab infection of leaves was significantly higher than in the integrated system, in case of the same cultivars, which means that the rate of decrease in leaf area was heavier in the organic production. Cultivars did not differ statistically in parameters of inner content when the integrated system was compared to the organic one. Results did not show statistical difference between loss of leaf area caused by apple scab and the parameters of inner content, which is attributed to the regeneration of leaf area due to favourable weather conditions. Data on apple scab and inner content of the fruit justify essentially the environmental systems of apple production.

 

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59-61.
Vol 17No 4-52011

The aim of this study was to test the in vitro sensitivity of two isolates of Monilinia laxa to fungicides approved in integrated and organic production systems. In vitro efficacy of 7 fungicides (Champion 50 WP, Kocide 2000, Nordox 75 WG, Olajos rézkén, Kumulus S, Rézkén, Rézoxiklorid) and another 6 fungicides (Score 25 EC, Efuzin 500 SC,... Systane, Folicur Solo, Zato Plusz, Rovral) approved in organic and integrated production systems, respectively, were tested against brown rot of sour cherry. The three isolates showed differences in sensitivity to fungicides. Fungicides (with active ingredients of copper and sulphur) applied in organic production showed relatively high percent growth capacity of M. laxa. Rézkén and Kocide 2000 showed the highest and Kumilus S the lowest efficacy against brown rot. Fungicides applied in integrated production showed relatively low percent growth capacity of M. laxa. Score 25 EC showed the lowest and Rovral and Folicur solso the highest efficacy against M. laxa.

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67
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57-61.
Vol 11No 12005

In a three-year study, the effect of two winter pruning treatments (strong and weak) on apple scab (Venturia inaequalis) and powdery mildew (Podosphaera leucotricha) was evaluated on six apple cultivars (`Rewena', `Elstar', 'Liberty', 'Gala', 'Pilot' and `Jonica') in two environmentally-benign apple production systems (integra...ted and organic). Results on apple scab showed that compared to the integrated production system, the pathogen caused significantly more infection on the leaves of cvs. `Elstar', 'Gala' and `Jonica' in the organic production system. The effect of pruning treatments on apple scab was mainly detectable on the cultivars susceptible to the disease such as cvs. 'Gala' and `Jonica'. The weak pruning correlated with a higher level of apple scab infection than the strong pruning, especially in the organic production system. Primary mildew infection was very low in both protection systems (below 2 %), therefore, effect of pruning on primary mildew infection could not be analysed. Low secondary mildew incidence was detected in the organically treated orchard part; however, incidence of powdery mildew was significantly higher (P> 0.05) on cvs. 'Elstar', 'Pilot' and `Jonica' in the integrated orchard part compared to cvs. 'Gala', 'Rewena' and 'Liberty'. The highest incidence of mildew was on cv. 'Elstar'. Pruning had very low effect on incidence of secondary mildew infection on leaves.

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31-33.
Vol 17No 4-52011

On the Horticultural Station of the Debrecen University, Pallag, two year old apple plantations of two varieties (‘Pinova’, ‘Golden Reinders’) have been studied, in autumn 2011. The growth of the cross section area of the trunk and the central axis were measured and compared to evaluate the effect of two alternative growing technologies...: integrated and/or ecological. ‘Pinova’ proved to grow more intensely than ‘Golden Reinders’ independently from the growing technology. The effect of the technology, however, was expressed in ‘Pinova’, where the integrated technology produced more vigorous growth than the ecological one. The effect of varieties on growing intensity was more distinct than the effect of growing technologies. Under the ecological growing system, the difference between the varieties studied is more accentuated regarding growing intensity. By that reason, in ecological growing plantations of varieties with moderate vigour ought to be stimulated by all possible techniques (as pruning, nutrition etc.).

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65
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71-74.
Vol 16No 12010

In a former paper we treated the same relation comparing varieties frown in the biological or organic system of growing, now the tests have been performed with samples grown by the integrated system. The scores registered properties as taste, skin, colour, consistency and size. In addition, we also explored the relation between general impressi...on and the individual properties.As first purpose, we started with collecting primary data on 15 samples taken from fruits grown by the integrated method and kept over 60–90 days in a store, then offered to the consumers. The test is based on an organoleptic assessment (records are registered in a questionary). The individual judgements are processed and coefficients of correlation between the traits (taste, skin, size, colour) calculated. The validity of the mathematically proved relations is considered to be decisive in judging the preferencial consumption of fruits.

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68
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65-67.
Vol 15No 42009

The two worldwide accepted environmentally freindly production systems are integrated and organic. The two systems shows considerably differences in several practical production technology elements. These effects come forward in generative processes (e.g. flowering, fruit setting) and vegetative features (e.g. growth). The aim of our study was ...to study the effect of pruning intensity on growth features of cultivars in integrated and organic apple production. In early spring of 2009, we determined two pruning intensities (strong and week) on six scab resistant and tolerant cultivars. Strong pruning resulted in stronger shoot growth for all cultivars compared to weak pruning. It is obvious that pruning intensity had more effect on vegetative performance than conditional differences originates from integrated and organic production.

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68
81
15-18.
Vol 16No 32010

Prima’, ‘Gala’, ‘Remo’, ‘Topáz’, ‘Idared’, ‘Releika’, ‘Resi’, ‘Rubinola’, ‘Rajka’, ‘Rewena’ and ‘Florina’ apple cultivars, both from organic and integrated farming, from Pallag and Újfehértó, were compared. Average size, weight, soluble solids, titratable acidity, total polyphenols, free radical sca...venging capacity expressed as Trolox equivalent (TEAC), copper and zinc were determined at harvest and after cool storage. Organic apples were more acidic, while integrated fruits had mostly higher copper and zinc content. Total polyphenols and TEAC values did not show a significant difference as a function of farming technology. A principal component analysis shows the separation of provenances as well as stored and fresh apples. Results are considered as preliminary.

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71
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57-60.
Vol 15No 42009

The aim our study was to establish whether significant differences in nutrients uptake and quality of soil and leaf exist between organic and integrated grown apples. The study was performed at the orchard Fruit Research Station, University of Debrecen, at Debrecen-Pallag during 2002–2004. Macro and micro elements were measured in soil and pl...ant samples. Analyses of variance of soil nitrogen data indicated highly significant differences between the two management systems (P < 0.001) for each examined nitrogen fraction. Analyses of variance of soil phosphate data indicated significant differences (P < 0.05) between the two management systems for orto-PO4 3– contents. Our data indicated that highly significant differences between the two management systems (P < 0.001) for magnesium, copper, and zinc; while significant differences between the two management systems was at P = 0.007 for calcium. Three year’s data of leaf phosphorus, sulphur and zinc were not shown significant differences between production systems. Nevertheless manganese and copper contents of leaves were higher in the organic orchard compared to the integrated one.

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76
86
11-14.
Vol 24No 3-42018

In our study, vegetative characteristics of 39 apple cultivars were evaluated in environmentally friendly production systems. Numbers of the branches of the central leader in different high zones were shown. According to our results, number of the branches of the axis was probably larger in the integrated production system, compared to the orga...nic one, which is related to the conditional status of the trees. Based on our experiences training and maintaining canopies in integrated system was easier, as relative more extensive canopies were needed in organic farming.

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119
100
35-39.
Vol 17No 4-52011

Summary: On the Experimental Farm of the Debrecen University at Pallag, a factorial experiment has been started at springtime of 2008 on an orchard of 12 year old apple trees ’Idared’ grafts on M 26 understocks. The first factor was the growing system (integrated versus ecological), the second was the intensity of pruning (strong versus mod...erate thinning of the crown), whereas the third was the method of pruning (shortening versus thinning of individual shoots). Measurements have been concentrated on shoot growth, flower initiation, fruit set and accumulated yield of two successive seasons. The following conclusions are made.
– In the ecological growing system, strong thinning proved to be beneficial for growth as well as for fruit bearing. – Strong thinning combined with shortening of shoots diminished yield in the first year, whereas
moderate thinning combined with shortening of shoots caused favourable growth and fruit bearing.
– In the ecological growing system, the moderate shoot growth was associated with better flower initiation, which increases the chances of alternate yielding.
– According to our results, the flower initiation of the ‘Idared’ was stimulated univocally by the thinning of shoots regardless of the intensity of pruning.
The right choice of the intensity and method of pruning is a decisive moment of a successful yield in the ecological apple growing system

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67
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113-122.
Vol 16No 12010

For adequate yields in apple plantations, during the long growing period of the fruit primordia, one of the decisive factors is water supply. Indicators of stress are valuable signs for the diagnosis of drought and necessity of watering, i.e. planning the irrigation of plantations. The aim of the present study was to find reliable signs of wate...r stress on apple trees and at the same time conclude on the drought tolerance of different varieties. The plantation of apple varieties grafted on various stocks and cultivated according to different systems (irrigated, non irrigated, integrate and biological) has been examined continuously by leaf analysis. Along the period of growing fruits, measurements were made in the field, then the leaf samples were analysed in the laboratory for composition of pigments, carbohydrates and antioxidants, as well as the histology of the tissues checked. Without irrigation, the mean leaf mass and the relative chlorophyll content (SPAD) of the variety Idared on M4 stocks increases beyond the values of 50 SPAD, whereas on M26 stock and integrated system, it declines. In the collection of varieties, grown according to the „integrated” system, during the development of fruits, ‘Gála’ and ‘Remo’ varieties have been affiliated to the “less susceptible” group regarding drought tolerance because of the leaf morphology, structure and content of SPAD and antioxidants. However, ‘Idared’ and ‘Jonagold’ belong to the “susceptible” group. To the same group are ranged the ‘Akane, Red Rome vanWell, Pink Lady’ varieties. Those varieties reacted to a short period of drought by increased production (content) of carbohydrates. Regarding changes of carbohydrate content ‘Greensleeves’ and ‘Ozark Gold’ varieties belong to the moderately water dependent (requiring) group. In the field, SPAD-tests facilitated the measurement of nutrient-uptake and incorporation, which is proved by the tight correlation between the data of SPAD and the increment of leaf weight (r=0.76–r=0.88), however, this depends on the variety too. SPAD is an indicator of water supply and is related with the density of stomata, cannot used for the selection of water-exigent varieties but for (drought) tolerant ones. In integrated culture, the (drought) susceptible varieties display (water-soluble) hydrophilic and lipophilic antioxidants (ACWand ACL resp.) the quantity of which may have some role in drought tolerance.

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82
100
11-14.
Vol 19No 3-42013

The indirect elements of the integrated plant protection can infl uence the effectiveness of the chemical control, however their significance are often extenuated. The characteristic of the plants are almost completely disregarded at the protection decisions. The condition can determinate basically the rate of the damages caused by the pests. T...he plant in good conditional state has higher tolerance and lower susceptibility against the pests. With the optimal general and annual condition we can establish and maintain the harmony between the growth and the bearing, what is the basis of effective plant protection.

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74
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37-40.
Vol 14No 1-2.2008

The year of 2007 was critical for fruit growers in the region of Easter-Hungary. Several orchards were suffered frost damages. Significant frost damage was also happened in the orchard of Tedej Rt., which caused total fruit failure in the orchard. Our study was conducted in the integrated apple orchard (Malus domestics Borkh.) establis...hed on a lowland chemozem soil in East-Hungary, to investigate how frost changes the mineral content of different apple species. Leaf samples were collected 100 days after full bloom both in 2006 and 2007. The concentrations of nitrogen, phosphorous, potassium, calcium and magnesium were measured in leaves. The absence of fruits caused a smaller accumulation of nitrogen and magnesium in leaves. Falling down of flowers and fruit sets hindered the translocation of the uptaken phosphorous, potassium and calcium towards fruit sets. Due to fruit failure the vegetative processes became dominant. Leaves larger amount of phosphorous, potassium and calcium stored. Besides the absolute element content, the ratio of the different elements was also determined. Majority of calculated ratios were removed from optimal values due to frost. Both absolute content of nutrients and their ratios pointed out that the frost damage significantly affected the uptake of nutrients and their storing processes.

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65
69
85-89.
Vol 16No 32010

The surface microbial contamination is of great interest in case of fruits, since they are they potential sources of spoilage or foodborne diseases. The aim of this work was to compare the epiphytic microbiota of food safety concern of different apple cultivars as a function of cultivation methods (organic versus integrated), production year an...d place. Investigating 47 samples it was found that the average microbial contamination of the apple surfaces was within a certain range,, practically independent of cultivar, growing area and year. The frequency distribution of the surface bacterial, mould and yeast counts did not show significant differences between growing technologies or varieties. The principal component analysis the samples ranked the apple samples into seven groups on the basis of their microbial contamination level, and the discriminant analysis proved the goodness of grouping. The grouping was independent from cultivars and growing methods. No foodborne pathogen Salmonella spp. and Listeria spp. were found on the surfaces of apples.

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69
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65-67.
Vol 19No 1-22013

The aim of this study was to evaluate the cherry leaf spot (CLS) incidence on 12 sweet cherry cultivars in integrated sweet cherry production in three consecutive years from 2010 till 2012. Assessed cultivars (cvs.) were ’Aida’, ’Axel’, ’Biggareau burlat’, ’Blaze Star’, ’Celeste’, ’Germersdorfi 3’, ’Izabella’, ’Kat...alin’, ’Krupnoplodnaja’, ’Linda’, ’Sunburst’, ’Vera’Results showed that year had an essential effect on the CLS disease incidence. Cultivars showed great differences within years. Cultivar ’Celeste’ showed the lowest disease incidence on leaves in all years lower than 10%. The largest disease incidence on leaves was on cv. ’Sunburst’ in 2010, on cv. ’Germersdorfi 3’in 2011 and on cv. ’Axel’ in 2012. Disease incidence on these cultivars ranged between 10 and 25%.

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76
90
11-14.
Vol 23No 1-42017

In this study effect of the environmentally friendly production systems on the brunch cross sectional area of apple trees trained to free spindle canopy are presented based on the data of 39 cultivars. According to our results branch thickness of the apple cultivars located in the lower, middle and upper region of the tree are more homogenous i...n the integrated production system compared to the organic one. Therefore the severity of the annual maintain pruning must be necessarily more moderate in the integrated growing system, than in the organic production system.

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143
102
15-18.
Vol 17No 4-52011

In modern and healthy diets antioxidants play an important role providing natural defence against serious diseases. Therefore it is recommended to include fruits and vegetables having high antioxidant capacity in daily diet in a due course. Apricot is one of the fruits receiving an increasing attention in this field. This study was conducted to... investigate the composition and content of fat-soluble carotenoids and tocopherols in different varieties of apricot using recently developed liquid chromatographic methods. Also it was aimed to compare organic and integrated farming in their effect on carotenoid and tocopherol content of the fruits. The results showed that apricot fruit are rich in vital carotenoids and bioactive tocopherols with significant variation between different varieties. The organic farming had favourable effect on the level of the major carotenoids and depending on variety this technology either increases or does not have significant influence on vitamin E content.

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