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• New data on the Hungarian adventive flora: Acer opalus Mill. subsp. obtusatum (Waldst. et Kit. ex Willd.) Gams 1925 in Hungary

  229-238

  Márton Korda

  Views:

  58

  Acer opalus Miller 1768 because of its complex taxonomic status can be interpreted as a collective species which contains several species and subspecies depending on different interpretations. An important adventive occurrence from this collective species was found in a forest that belongs to the Tétényi plateau (Budapest, 11th district) in July of 2009. It is proved to be Acer opalus Mill. subsp. obtusatum (Waldst. et Kit. ex Wild.) Gams. The natural range of the subspecies covers the eastern side of the central Mediterranean Sea region and western Balkan Peninsula. It is a submediterranean-mediterranean, thermophile, shade-tolerant taxon but often appears together lightdemanding species too. In its native range it occurs in beech, pine, sessile oak forests and in different types of dry oak forests as admixed tree species. The subspecies was found in a turkey oak – sessile oak community in Hungary. Several old, seed growing individuals live on the spot which have had continuous seed production. Therefore from the old trees to the young seedlings different age categories are represented.

• Rediscovery of Orchis mascula subsp. signifera (Vest) Soó in the Keszthely Mts (Transdanubia, Hungary)

  51–54

  Miklós Óvári

  Views:

  143

  Orchis mascula subsp. signifera is a rare, legally protected and endangered species of the Hungarian flora. Currently it is recorded from 25 flora mapping grid cell in Hungary. The westernmost Hungarian occurrence of the species was documented by a voucher collected by Dénes Gotthárd in 1976. After four decades the occurrence of O. mascula in the Keszthely Mts was confirmed in 2015. The local habitat preference and major threats to the taxon are briefly discussed.

• Data to the flora and vegetation of Hungary III.

  197-206

  László Erdős

  Eszter Aradi

  Zoltán Bátori

  Csaba Tölgyesi

  Views:

  149

  We report the occurrence data of 44 vascular plant taxa collected at various localities in Hungary (mainly the Danube–Tisza Interfluve and South Transdanubia) during floristic, vegetation and ecological field surveys between 2016 and 2018. Of the species in the present article, 27 are protected or strictly protected. In the paper we focused on plants that are rare regionally (e.g. Botrychium lunaria, Polystichum aculeatum) or in the whole country (e.g. Ophrys oestrifera, Ranunculus psilostachys), as well as on plants that occurred in unusual habitats (e.g. Corydalis solida). We also included new localities of weedy species, some of which (e.g. Opuntia humifusa, Sporobolus cryptandrus) may present nature conservation threats.

• Contributions to the Atlas Florae Hungariae V.

  383–403

  János Csiky

  Kornél Baráth

  Virág Bocz

  Judit Deme

  Zsófia Fülöp

  Dániel Kovács

  Katalin Nagy

  Bendegúz Tamási

  Éva Csikyné Radnai

  Views:

  167

  The current paper is the 5th one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Several distribution data of 360 plant taxa from 66 flora mapping units are published in this study. Most of them are from the South Transdanubia. As a result of systematic surveys, more than 30 new species were found in 6 flora mapping quadrates. Most of the species are frequent, treated usually as weeds, but some of them are rare (e.g. Anchusa azurea, Centunculus minimus, Euphorbia stricta, Hypericum humifusum, Sagina apetala), and in some cases they are legally protected species in Hungary (e.g. Asplenium adiantum-nigrum, Dryopteris affinis, Festuca dalmatica, Gymnocarpium dryopteris, Urtica kioviensis).

• Contributions to the flora of the Szuha watershed and adjacent territories (NE Hungary)

  27-67

  Péter Szentgyörgyi

  Gábor Bátori

  Views:

  289

  This paper presents new floristic data of 415 taxa from the Putnok Hills and Sajó Valley microregions (Northern Hungary). Records were collected between 1991 and 2021. In the first part new occurrences of legally protected (e.g., Cypripedium calceolus, Iris aphylla subsp. hungarica, Lycopodium clavatum, Thalictrum aquilegiifolium, Pyrola rotundifolia, Scorzonera purpurea, Stipa pulcherrima) and not protected but locally rare or data-deficient species (e.g., Myosurus minimus, Thalictrum simplex, Sisymbrium loeselii, Camelina microcarpa, Potentilla collina, Trifolium striatum, Geranium pratense, G. dissectum, G. divaricatum, Viola pumila, Epilobium roseum, Myriophyllum spicatum, Pimpinella major, Berula erecta, Trinia ramosissima, Centaurium pulchellum, Nepeta cataria, Limosella aquatica, Veronica scutellata, Orobanche arenaria, Utricularia vulgaris, Galium elongatum, Cephalaria transsylvanica, Dipsacus fullonum, Xanthium saccharatum, Alisma lanceolatum, Elodea canadensis, Potamogeton pectinatus, P. berchtoldii, P. lucens, P. nodosus, Zannichellia palustris, Sisyrinchium bermudiana, Melica picta, Elymus elongatus, Leersia oryzoides, Typha laxmannii, Eriochloa villosa) are presented. This chapter contains data on 246 species from which 8 are new for the flora of the Tornense flora region and 47 for its subregion, the Putnok Hills. Further contributions to the distribution data of Atlas Florae Hungariae are presented in Electronic Appendix. This section contains raw occurrence data of 169 species.

• Epipactis exilis in the Börzsöny Mts (C Hungary)

  102-103

  Balázs Pintér

  Gábor Tímár

  Views:

  208

  Seven specimens of Epipactis exilis has been found in the Börzsöny Mts, in a beech forest along the creek Szén-patak on the 21st August 2020. This strictly protected species is very rare in Hungary, the present observation is its fifth occurrence in the country. It is new to the Börzsöny Mts flora.

• Interesting moss and liverwort records from the Western Mecsek Mts (South Transdanubia, Hungary)

  29-38

  János Csiky

  Bianka Atkári

  Judit Deme

  Éva Csikyné Radnai

  Views:

  84

  During an extensive field work in the acidophilous forests and fens of the Western Mecsek Mts. current localities of several species, which haven’t been recognised here for 50 years, were revealed (e.g. Bazzania trilobata, Diphyscium foliosum, Scapania nemorea). The authors also found some new stands of regionally and/or locally rare taxa (e.g. Buxbaumia aphylla, Dicranum tauricum, Jamesoniella autumnalis, Lepidozia reptans, Sphagnum squarrosum) as well as species new to the moss flora of the Mecsek Mts. (e.g. Aulacomnium androgynum, A. palustre, Calliergon cordifolium, Hedwigia stellata). Both investigated habitat types were found to be infected by invasive moss (Campylopus introflexus) and liverwort (Riccia rhenana) species. It is noteworthy that most of the species were found in winter or at the beginning of spring, when most vascular plants are inactive and their coverage is negligible. In this period moss and liverwort taxa are active, and are therefore easily noticeable. The authors raise the possibility that the expansion of Dicranum tauricum in Hungary is a consequence of former coniferous (especially Pinus spp.) plantations, since the old, dead trunks of pine species can serve as stepping stones to colonization for this taxon.

• Data to the flora and geobotany of Kisalföld (Lesser Plain) region, NW Hungary

  235-253

  Gergely Király

  Gábor Takács

  Angéla Király

  Views:

  128

  Lesser Plain (Kisalföld) – divided among three countries (Austria, Hungary and Slovakia) – is the westernmost part of the lowland areas of the Pannonian Basin. Present paper provides a résumé of the authors’ floristic work from the Hungarian part of the area, describing localities, habitats and regional phytogeography of 57 taxa altogether. The localities were listed according to the classification of small geographic regions. The most important results of the study are as follows:

  • We recorded 5 species new to the region (Brachypodium rupestre, Bromus ramosus, Hypericum dubium, Lathyrus sphaericus, Tordylium maximum); all of them can be considered as submontane-montane elements.
  • We reported several species new to a microregion within the Lesser Plain: montane species in the W-SW part of the Plain (Cirsium rivulare, Dryopteris dilatata, Equisetum telmateia), loess-connected species mainly on the Moson Plateau (Agropyron cristatum, Allium atroviolaceum, Euphorbia salicifolia, Viola ambigua); and occurrences of species on sandy soils near the Lake Fertő (Draba nemorosa, Stipa pennata).
  • We explained the known localities of several salt tolerant species (e.g. Carex divisa, Crypsis alopecuroides, Juncus maritimus, Limonium gmelinii, Suaeda pannonica) from the Lake Fertő and its surroundings.
  • We assessed the role of the forest management and nature conservational actions in the distribution of some important tree species (Acer tataricum, Betula pubescens, Fraxinus ornus, Quercus petraea, Salix pentandra).
  • We discussed the regional nature conservational status of several protected and/or threatened species (e.g. Allium carinatum, Allium suaveolens, Carex strigosa, Hottonia palustris, Potamogeton coloratus, Stellaria palustris) in the Lesser Plain.

  The activities of the authors in the surroundings of Győr were supported by monitoring results gained during the Hungarian Little Plain project (LIFE08 NAT/H/000289).

• Remnants of closed oak woods on loess in the Mezőföld (Pulmonario mollis-Quercetum roboris Kevey 2008)

  66–93

  Balázs Kevey

  András Horváth

  Gábor Lendvai

  Views:

  159

  The phytosociological characterization of closed oak forests occurring in the loess-covered part of Central Hungary (Mezőföld) is presented. Such forest fragments are found very sporadically in that predominantly treeless region, mostly in semi-dry habitats of north-facing hillsides. This plant community represents an intermediate stage between open steppe woodland (Aceri tatarici-Quercetum pubescentis-roboris) and closed mesic oak-hornbeam forests (Corydali cavae-Carpinetum). Analyses of 20 phyto­sociological samples using multivariate grouping methods (cluster analysis, principal coordinates analysis) showed a clear difference between these samples and samples of similar communities occurring in the area. In the species composition the characteristic elements of dry and mesic oak forests (Quercetea pubescentis-petraeae, Quercetalia cerridis, Aceri tatarici-Quercion and Fagetalia, respectively) played a major role. On the other hand, dry grassland species (Festuco-Brometea, Festucetalia valesiacae, Festucion rupicolae, etc.) were much less significant than in steppe woodlands. In terms of chorology, European and sub-Mediterranean floristic elements were dominant in the samples, while the proportion of continental elements was substantially smaller than that in steppe woodlands. As a result, our samples were identified with the plant association Pulmonario mollis-Quercetum roboris Kevey 2008, which is classified into the suballiance Polygonato latifolio-Quercenion roboris Kevey 2008 in the phytosociological system.

• Epipactis pseudopurpurata Mered’a, a new orchid species for the Hungarian flora

  27–32

  Miklós Csábi

  Antal Halász

  Views:

  99

  A small population of Epipactis pseudopurpurata new to the flora of Hungary was found in the Buda Mts, near Telki village (Pest county). The main diagnostic features of this species are discussed, and a comparison with the allied species, E. purpurata is provided.

• Taxiphyllum densifolium (Taxiphyllaceae) and other rare bryophytes of the Mecsek valleys (SW Hungary)

  3-16

  János Csiky

  Attila Sipos

  Views:

  88

  In this paper, we deal with threatened and/or legally protected bryophytes found in the shady, forested valleys of the Mecsek Mountains, which are rare in national and/or local terms. Taxip­hyllum den­sifolium, a rare moss in Europe has stable colonies with sporophytes in two valleys. This study is the first to describe the spore characteristics of this species. The previously considered unique and legally pro­tected Rhynchostegiella teneriffae is nowadays so widespread in the Mecsek that it can no longer be consi­dered a vulnerable species (VU) in Hungary. We present the first population of the legally pro­tected Nec­kera pennata in Tolna county, and we also report on the recent, unusual occurren­ce of the data-deficient (DD) Fossombronia wondraczekii. We also list new populations of three other endangered (EN) species, Blindiadelphus recurvatus, Palustriella commutata and Porella arboris-vitae. Several vulnerable (VU) taxa such as Blepharostoma trichophyllum, Porella cordaeana, Sciuro-hypnum flotowianum, Seligeria pusilla, as well as new occurrences of near threatened (NT) mosses such as Mic­roeurhynchium pumilum (with sporophytes), Phaeoceros carolinianus are documented. In addition, the data on Rhynchostegium rotundi­folium in Jakab Mountain, which has not been recorded for more than 50 years, is confirmed. The difficul­ties of differentiating Hygroamblystegium species are briefly discus­sed.

• Contributions to the Atlas Florae Hungariae III.

  122-146

  Csaba Molnár

  Győző Haszonits

  Ákos Malatinszky

  Gergely Károly Kovács

  Gábor Kovács

  Timea Nagy

  Attila Molnár V.

  Attila Takács

  Views:

  296

  The current paper is the third one in the series aiming to contribute with new distribution data to the distribution maps published recently in Atlas Florae Hungariae. Current occurrence data of 558 vascular plant taxa from 186 flora mapping quadrates (CEU) are presented. New records are indicated from different regions of Hungary; however, most occurrences are located in the North Hungarian Mts. Occurrence data of rare native taxa (e.g. Marsilea quadrifolia, Salicornia prostrata, Stellaria palustris, Potentilla patula, Althaea cannabina, Lythrum tribracteatum, Ajuga laxmannii, Plantago schwarzenbergiana, Alisma gramineum, Gagea bohemica, G. szovitsii), rare or data-deficient alien taxa (e.g. Chorispora tenella, Thladiantha dubia, Senecio vernalis) as well as frequent but more or less underrepresented taxa (e.g. Scleranthus annuus, Vicia hirsuta, V. lathyroides) are also enumerated. In case of the most common species we provided the CEU codes only.

• Floristic records from Northern Hungary II. North Cserehát and Upper Bódva valley

  31–48

  András Vojtkó

  Tünde Farkas

  Views:

  291

  This paper presents new floristic data of 123 taxa from the regions of North Cserehát and Upper Bódva valley (Northern Hungary). Records were collected between 1995 and 2020. New occurrences of legally protected (e.g., Dactylorhiza incarnata, Eriophorum angustifolium, Lycopodium clavatum) and not protected but locally rare or data-deficient species (e.g., Artemisia annua, Carex remota, Cirsium pannonicum, Cyperus fuscus, Genista pilosa, Melampyrum cristatum Ranunculus flammula, Scorzonera purpurea, Silene noctiflora) are presented.

• Floristic data from the central part of the floristic region ‘Crisicum’ (E Hungary)

  317–357

  Balázs András Lukács

  Gergely Gulyás

  Dénes Horváth

  István Hődör

  András Schmotzer

  Gábor Sramkó

  Attila Takács

  Attila Molnár

  Views:

  249

  This paper reports new floristic data of 207 taxa. Data collected between 2005 and 2015; originated from the central part of the floristic region ‘Crisicum’ (i.e. the E part of the Great Hungarian Plain), which is a currently poorly studied area of Hungary from a floristic point of view. The paper reports occurrence localities for some rarities such as Asplenium trichomanes, Marsilea quadrifolia, Sisymbrium polymorphum, Viola stagnina, Gentiana pneumonanthe, Elymus elongatus, Montia fontana subsp. chondrosperma, Ranunculus rionii, Ranunculus illyricus, Myagrum perfoliatum, Sedum caespitosum, Vicia biennis, Dorycnium herbaceum, Oenanthe banatica, Peucedanum cervaria, Echium italicum, Digitalis lanata, Valerianella rimosa, Craex buekii, Cephalaria transsylvanica, Cyperus pannonicus.

• New montane species in the Hungarian bryophyte flora

  16-26

  Csaba Németh

  András Schmotzer

  Views:

  652

  During a bryofloristical investigation in the vicinity of Ómassa (Miskolc) in the Bükk Mountains (NE Hungary), three montane-boreal bryophyte species new to the Hungarian bryoflora were collected. Encalypta spathulata Müll.Hal. was found on the ledges of limestone cliffs of Vörös-kő Hill as well as on the rocky debris among roots of a fallen tree lying under the cliffs. Hydrogonium croceum (Brid.) Jan Kučera and Orthothecium rufescens (Dicks. ex Brid.) Schimp. were found in the crevices of dolomite cliffs on the north-facing steep slope of Jávor Hill. While the latter two species are widespread in European Mountains, E. spathulata is considered to be a rare species, and is assessed as ‘Vulnerable’ in the IUCN red list.

• Some floristic data from surroundings of Kötcse (South Transdanubia, West Outer Somogy)

  74-80

  Timea Nagy

  Views:

  99

  There are very few published and documented floristic data from the hills of the SW region of Hungary called 'Outer Somogy'. This paper present the occurrence data of 31 vascular plant species from the western part of this region. Among them 14 taxa are legally protected and further 17 taxa are sporadically occurring. The records were collected within the administrative area of villages Karád, Kereki, Kötcse, Nagycsepely, Szólád (Nezde) and Teleki.  Most data were recorded in 2014. This is the first report on the occurrence of 1) Neotinea ustulata subsp. aestivalis in South Transdanubia, and 2) Epipactis microphylla and E. tallosii in Outer Somogy. Data provided on the occurrence of Cirsium boujartii on the northwestern border of the distribution area of this insufficiently known species.

• Bryofloristical study in the Brunszvik manor park in Martonvásár, Hungary

  198–206

  Zoltán Nagy

  Imre Majláth

  Marcell Molnár

  Péter Erzberger

  Views:

  153

  This is the first bryofloristic study in the Martonvásár manor park. In our survey in 2015, 56 species (6 liverworts and 50 mosses) were found in the park. Most of them are considered to be common in Hungary, but an endangered (Eurhynchium speciosum), a vulnerable (Orthotrichum patens) and five near-threatened species (Aphanorrhegma patens, Orthotrichum obtusifolium, Orthotrichum pumilum, Pseudocrossidium revolutum, Riccia cavernosa) were also found. These results point out the nature conservation value of the manor park.

• New data to spreading of Plantago coronopus in Hungary

  99-101

  Dávid Schmidt

  Views:

  262

  In this study, new data of the naturalized roadside alien Plantago coronopus L. found in 2020 are listed. Numerous stands have been detected at sites far from previously known localities. These data show that P. coronopus is still in an intense phase of its spread. The species currently known from 90 flora mapping units in Hungary. Its new observations come from the narrow vegetation strip along roadsides, with the exception of the new data from Szombathely, which comes from the parking lot of a shopping center.

• Campylopus flexuosus (Hedw.) Brid.: a moss new to the Hungarian bryophyte flora

  22-28

  Péter Erzberger

  Csaba Németh

  Views:

  99

  Two localities of Campylopus flexuosus new to Hungary were discovered by the authors near Uzsa in the Bakony Mts during mapping of the Hungarian bryoflora. In this paper the circumstances of the occurrences are described and illustrated by maps and photographs; a short description of the species is provided together with photographs of diagnostic characters.

• Multiclavula mucida (Basidiomycota) and other cryptogamic taxa in the Hungarian flora

  173–184

  János Csiky

  Judit Deme

  Peter Erzberger

  Dániel Kovács

  Csaba Németh

  Attila Sipos

  Views:

  220

  The paper deals with the species found during cryptogamic flora mapping in 2023 that are remarkable from a floristic, taxonomic or conservational point of view. We present data from the Bükk Mts of two Trapeliopsis species which appear to be rare in Hungary. Multiclavula mucida, which is ap­parently a rare basidiomycete lichen species in Europe that prefers humid, montane habitats was dis­covered in the Mecsek Mts as a new species for the Hungarian flora. Of the knothole moss, Anacampto­don splachnoides, which is now thought to be entomophilic, we communicate the only extant popula­tion from Southern Transdanubia. We report the first occurrence of the atlantic-mediterranean Lepto­don smithii from the Hungarian side of the Great Hungarian Plain. Numerous data of the corticole Neck­era pennata indicate that in Hungary the distributional centre of this species is situated in the county of Baranya in the lowlands. New data from the Bükk Mts underline that in the Pannonian region Buxbaumia viridis is primarily a terricole species of acidophilous beech forests. We communicate the first occurrence from the lowlands of the alien lignicolous Sematophyllum adnatum which is danger­ous­ly spreading in Europe, at the same time one of the most vigorous populations in the Pannonian region. The third Hungarian occurrence of Callicladium haldanianum, a species that seems to spread in bogs, as well as the second occurrence of Racomitrium lanuginosum, growing abundantly on an ande­site boulder scree in Mátra Mts, are also reported here. Of Phegopteris connectilis we present in photo­graphic docu­mentation the first extant population from Southern Transdanubia, and we discuss whether the first publication from Mecsek Mts is correct or exact. In addition to listing the floristic data, we also briefly discuss the (mis)use of distribution data in the context of climate change, the difficulties of distinguish­ing the microtaxa of some pteridophytes, e.g. Asplenium adiantum-nigrum agg. and Dryop­teris affinis agg., and we correct one of our previously published, erroneous Hedwigia stellata data. Some interest­ing data of other taxa (such as Leucobryum glaucum, Palustriella commutata, Tetraphis pellucida, Gym­nocarpium robertianum, Ophioglossum vulgatum) are also mentioned in the paper.

• The occurrence of Pterygoneurum squamosum Segarra et Kürschner on a reclaimed waste storage near town Győr (NW Hungary)

  23-27

  Péter Szűcs

  Views:

  107

  In the article a new locality, range map and habitat description of Pterygoneurum squamosum is provided from Hungary. Differently from up to the present known localities, the moss was found on anthropogenic habitat, eroded and clayey soil surface. Considering the climatic and soil conditions there, probably this rare Mediterranean species has wider ecological tolerance, so we come across it on further man-made habitats, as well as on loess walls.

• Some floristic data from surroundings of Kötcse II. (West Outer Somogy, South Transdanubia, Hungary)

  114-121

  Timea Nagy

  Views:

  95

  Floristic data of 73 species from NE part of West Outer Somogy are provided in the paper. Most of them were collected in 2016. The paper contains distribution data of species, which have not been detected in Outer Somogy so far (e.g. Peucedanum arenarium), and new occurrences of rare or sporadic taxa (e.g. Menyanthes trifoliata, Veronica scardica), as well as subspontaneous occurrences of cultivated species (e.g. Narcissus poëticus, Helleborus purpurascens) are provided.

• Current distribution of the potentially invasive narrow-leaved ragwort (Senecio inaequidens DC.) in Hungary

  179-187

  Győző Haszonits

  Dávid Schmidt

  Views:

  227

  Our study presents the actual status of potentially invasive Senecio inaequidens in Hungary. Literary and herbarium data are also discussed. We report some new data from ruderal places and near highways which are unmarked until now. The actualized distributional map of the species are given. We sign the potential habitat threats too.

• Additions to the flora of Jászság (Central Hungary)

  21–30

  Csaba Molnár

  Views:

  295

  Within the framework of the Hungarian Flora Mapping Program, nine quadrants were surveyed in the territory of Jászság in 2012 and 2020. Regionally important floristic records of 68 vascular plant species are presented in this paper. Some of them are aliens (e.g., Alopecurus myosuroides, Amaranthus deflexus, Juncus tenuis, Phytolacca esculenta) or rare native weeds (e.g., Aegilops cylindrica, Bupleurum affine, Myagrum perfoliatum). Records of regionally (e.g., Coronopus squamatus, Linaria biebersteinii) or nationwide (Silene bupleuroides) rare native species as well as indicators of former forests (e.g., Clematis recta, Elymus caninus, Melica altissima, Sisymbrium strictissimum) are also presented. An interesting road edge rich in saline species (Artemisia santonicum, Chenopodium chenopodioides, Hordeum hystrix, Plantago maritima) is discussed in detail.

• The role of soil seed bank in restoration and dynamics of Hungarian plant communities

  116-135

  Réka Kiss

  Views:

  118

  Soil seed bank has an important role in maintaining plant populations and communities, in regeneration processes, and also provides information about the history of the studied site, the past land use practices, the biotic and abiotic changes and about the current degradation degree. In Hungary there are many researchers related to the seed bank, but from the last few years we do not have a comprehensive work about the results of these works. My goal is to report on the results of former seed bank researches in Hungary and also to highlight the missing topics and further research opportunities. The review is based on 49 articles, the oldest one dates back to 1922, the newest one was published in 2015. Most of the studies use greenhouse or laboratory germination tests to determine the viability of seeds. The earliest studies, which examined the weed seed bank of farmland soils found, that the seed bank was much higher than the total seed bank found on natural and semi-natural habitats studied later. Researchers found the lowest seed bank density in forests. Disturbed sites and former agricultural sites had higher seed bank densities than sites without agricultural use. Most part of seed bank was located in the upper soil layer in all studied communities. In most studies usually a small similarity was detected between the composition of soil seed bank and aboveground vegetation, which suggests that the seed bank has a subordinate role in regeneration processes. Seed bank type classification is lacking from the three-quarter of Hungarian flora species, despite that it is essential for protecting species and communities and for developing proper protection against invasive and alien species.