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The herbarium of Debrecen University (DE) I. The “Rezső Soó Herbarium”
142-155Views:130The paper introduces the herbarium of Debrecen University (DE). All specimens of the herbarium were documented by digital photographs, and all data from the labels were entered into an Excel spreadsheet. The largest collection of Herbarium DE, the “Soó Rezső Herbarium” (abbreviation: SRH) consists of ca. 38.000 specimens from almost all European countries, but mainly from Hungary and the neighbouring countries. Most of the specimens originate from Hajdú-Bihar, Pest, Győr-Moson-Sopron, Borsod-Abaúj-Zemplén, Veszprém and Szabolcs-Szatmár-Bereg counties. The SRH stores a considerable amount of original/authentic specimens collected by Hungarian botanists, primarily by Rezső Soó and his followers, as well as Sándor Polgár. The most rapid growth of the SRH took place in the first half of the last century (1930–1950). By contrast, the collection was practically not developed between 1955 and 2000. After the millennium it started to grow again. Data on the most substantial part of the SRH covering native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting date, collector, file name of documentary photograph).
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Herbarium database of the vascular collection of Eszterházy Károly College (EGR)
339-348Views:118The paper describes the vascular herbarium of the Eszterházy Károly College (EGR) in Eger (Hungary), according to its condition in 2013. All specimens of the herbarium were documented by digital photographs (ca. 8 000 specimens), and all data from the labels were entered into MS Excel spreadsheet. 54% of the specimens were collected in present-day Hungary, the other half comes mainly from the neighbouring countries, but more distant European countries are represented as well. Hungarian specimens were collected mostly in Heves, Pest, Zala, Vas, Borsod-Abaúj-Zemplén and Veszprém counties. Most of the herbarium sheets originated from the 1860’s and the 1950–60’s. Apart from these periods the collection has hardly developed. The most prolific collectors were Gábor Vida, Márton Vrabélyi, Árpád Károlyi and Tamás Pócs. More than half of the Hungarian flora (61%) is represented in EGR, although some taxa (e.g. Pterydophyta, Gymnospermatophyta) are much underrepresented. The digital photographs and the database are property of the authors and the Department of Botany and Ecology of Eszterházy Károly College. Data of non-cultivated plants collected in the territory of present-day Hungary are summarized in an electronic appendix (http://kitaibelia.unideb.hu/?download&aid=852&volume_id=94&lang=hun – including: catalogue number, taxon name, collector, settlement, date of gathering and file name of the documentary photograph). Further data can be required from the corresponding author or the curator of the herbarium.
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The Herbarium of Debrecen University (DE) II.: The „Zoltán Siroki Herbarium”
15-22Views:364The paper introduces the “Zoltán Siroki Herbarium” (on the basis of the MS Excel spreadsheet database of the collection) which is a separate section within the Herbarium of Debrecen University (DE). The Zoltán Siroki Herbarium consists of ca. 20 000 specimens from almost all European countries, but mainly from the historical territory of Hungary. Although ca. 400 botanists contributed to this collection, most of the specimens were collected by Zoltán Siroki in Hajdú-Bihar, Pest and Borsod-Abaúj-Zemplén counties. The most rapid growth of the Herbarium took place in the middle of the last century (from the 1940s to 1960s), but Siroki remained an active collector until the 1980s. Some specimens from the Herbarium of former Royal Seed Testing Station (Budapest) was also incorporated in this collection; they represent gatherings from the last decades of the 19th century and first decades of the 20th century. Data on the most significant part of the collection, representing native plants collected in the present-day Hungary, are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).
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In memoriam Zoltán Siroki (1906–1987)
3-14Views:92Zoltán Siroki (1906–1987) was a Hungarian agronomist and lecturer. His professional activity was connected mainly to Debrecen. Beside his widely known ornithological studies his floristic and fitocoenological data to Matricum, Crisicum and Nyírségense Hungarian regions are also important. Herbarium of Zoltán Siroki, consisting of ca. 20 000 specimens is a remarkable part of the Herbarium of Debrecen University (DE).
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The herbarium of the Botanical Garden of Eötvös Loránd University (BPU)
55–59Views:172The paper introduces the herbarium of Eötvös Loránd University (BPU), currently stored at the Botanical Garden of the University, according to its state in the year 2013. The BPU herbarium consists of ca. 16 000 specimens the relevant data of which have been organised into an OpenOffice Calc spreadsheet database. The specimens were collected in 16 European countries, mainly in the current territory of Hungary (80%), Romania (12%) and Slovakia (5%). Other countries are represented with very few specimens. The vast majority (ca. 70%) of the Hungarian specimens were collected in Pest, Veszprém, Borsod-Abaúj-Zemplén, Hajdú-Bihar and Szabolcs-Szatmár-Bereg counties. Although the gatherings cover a century and a half period of time, the most rapid growth of the collection took place from the 1930s to the 1960s. The most prolific collectors were Rezső Soó and his followers: Lajos Felföldy, Tibor Simon and Szaniszló Priszter. Felföldy and Simon enriched the collection through more than 60 years. Data on the native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).
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On the former occurrence of Calla palustris in Hungary
200-210Views:1674The origin of a more than 150 years old herbarium specimen of Calla palustris L. stored in JPU is reconsidered in this study. According to the information on its label, the specimen was collected in 1856 at Lake Velence, close to Székesfehérvár city (central Hungary). Some Hungarian botanists are skeptical about the authenticity of this record. Based on the current occurrences of some remarkable swamp species (e.g. Carex pseudocyperus, Liparis loeselii, Sphagnum spp.) and habitat types (e.g. willow carrs and swamp forests) at the lake, as well as the habitat preference of bog arum, there is no reason to question the former occurrence of the species here or even in other Hungarian localities. The content and accuracy of the examined voucher meet the expectations of floristic data. However, in the absence of recent data the species can be considered extinct from Hungary.
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Dr. Sándor Polgár was born 140 years ago
169–184Views:112Dr. Sándor Polgár was the most outstanding botanist of Győr county (NW Hungary). He provided important results in floristic studies, phytogeography, taxonomy and in the research of alien plants. „Győr megye flórája” („Flora of Győr county”), published in 1941, was one of the most important monography in his period. He is the author of the rare, hybridogenous species Ornithogalum ×degenianum, known only from Hungary. His private herbarium was one of the biggest in Hungary with more than 20,000 specimens. As a teacher he taught geography and nature studies in his home town Győr between 1900 and 1935. Because of his Jewish origin, he was a victim of the holocaust in 1944.
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Dr. Antal Waisbecker was born 180 years ago
179–192Views:130Dr. Antal Waisbecker (1835–1916) was a Hungarian physician and the most prominent botanist of Vas county (West Hungary) at the turn of the 19–20th century. He contributed with important floristic data for the botanical monography of Vas county, authored by Vince Borbás. He processed the pteridophytes of Vas county and cormophytes in the surroundings of Kőszeg. Antal Waisbecker was an expert in the knowledge of sedges too. He discovered and described the Carex fritschii. Remarkable amount of herbarium specimens collected by Waisbecker are deposited at the Savaria Museum (Szombathely, SAMU) and Hungarian Natural History Museum (Budapest, BP).
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Data on the occurrence of sedge species (Carex, Cyperaceae) in and around South Nyírség (East Hungary)
165-198Views:381We provide floristic data on a total of 38 sedge species in the area of South Nyírség and its immediate surroundings. We also detected some populations of previously undescribed or rare species in the Hungarian Great Plain, such as Carex buxbaumii, Carex hartmanii, Carex umbrosa, and Carex paniculata. We report several new occurrences of species with little literature data from the area, such as Carex brizoides, Carex michelii, Carex pallescens, Carex elongata. However, during the last 13 years, we were not able to detect seven species previously mentioned in the literature. Herbarium data confirmed that some of the older Carex nigra occurrences for the area actually refer to Carex cespitosa. Based on herbarium data and the literature, we think that local data on Carex pairaei may largely refer to Carex divulsa subsp. leersii and/or Carex spicata. Contrary to many pieces of literature, Carex vulpina was not observed inside the sandy area.
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The ‘rediscovery’ of the Birdsfoot Fenugreek (Trifolium ornithopodioides (L.) SM.) in the Hortobágy, Hungary
207–212Views:287In 2009, the first author discovered an unknown population of Trifolium ornithopodioides in the Ágota-puszta of Hortobágy plain. In 2016, the second author found an another population in the vicinity of Karcag (Ecse-zug puszta). Up to now there was only one published occurrence of this species from the region. The specimen collected by Szujkó-Lacza Júlia, Kováts Dezső and Fekete Gábor in 1974 was deposited in the herbarium of the Hungarian Natural History Museum (BP), but this incomplete specimen was misidentified, and not Trifolium ornithopodioides. Therefore, this newly discovered site at Ágota-puszta should be regarded as the first trustworthy occurrence of this species at the Hortobágy plain.
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Sándor Polgár and the research of the adventive flora of Hungary
188–197Views:93In the 20th century Sándor Polgár was one of the most outstanding researchers of adventive plant species in Hungary. He wrote nine publications in this theme. In the industrial environment of his home town Győr (NW Hungary) he found tropical species, most of them were southern-American origin. The richest alien flora came around the oil factories of the town. He reported 65 taxa new for the Hungarian flora, four of them were new for Europe too. He was a great expert of problematic Solanum, Amaranthus and Chenopodium genera.
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Contributions to the lichen flora of the Mecsek Mts (South Hungary)
16–26Views:65A checklist of 115 lichen taxa was compiled from the Mecsek Mts, based on floristical records of 221 old or recently collected herbarium specimens (incl. 37 duplicates) deposited in the JPU herbarium (Pécs, Hungary). The occurrence of 22 taxa are new for the Mecsek Mts: Arthopyrenia cinereopruinosa, Baeomyces rufus, Candelariella reflexa, Catapyrenium rufescens, Cladonia macilenta, Dimerella pineti, Hypogymnia tubulosa, Ochrolechia arborea, Pleurosticta acetabulum, Pseudevernia furfuracea, Psilolechia lucida, Pycnothelia papillaria, Rinodina sophodes, Tomasellia arthonioides, Toninia candida, Trapelia involuta, Varicellaria hemisphaerica, Verrucaria marmorea, V. nigrescens, V. parmigera, Xanthoparmelia conspersa, X. verruculifera. One legally protected lichen species (Peltigera leucophlebia) is also represented by two specimens.
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Lajos Felföldy: a prominent Hungarian botanist and hydrobiologist
3–25Views:126Lajos Felföldy (1920–2016) was one of the most versatile and open-minded Hungarian biologists. He began his scientific career as a student of Prof. Rezső Soó. Between 1938 and 1946 he participated in geobotanical studies in University of Debrecen and Kolozsvár (Cluj-Napoca, Romania). He contributed with important achievements to the development of several biological disciplines. His pioneering study on the effects of air pollution on epiphytic lichens (1942) was among the firsts in the world. He described Hemitherophyte life-form as a discrete unit within Raunkiaer's plant life-form system (1942). He was a pioneer in the cytological (caryological) study of wild vascular plant species in Hungary (1947–1949). His results regarding to primary production of freshwater algae and algal culture (1958–1960) were in leading edge. In 1972 he founded and until 1990 edited the Hungarian series entitled ‘Vízűgyi Hidrobiológia’. Books of this series aimed to publish identification keys of freshwater taxa. These books were proved to be decisive and useful tools for Hungarian hydrobiologists in biological classification of brooks, streams, rivers and different types of stagnant waters. Between 1934 and 2009 he collected more than ten thousand herbarium sheets. After his retirement, he dealt with the revision of the herbarium material of Department of Botany in Hungarian Natural History Museum and Botanical Garden of Eötvös University (Budapest). He participated in the preparation of the New Hungarian Herbal. His scientific work was characterised by deep humility for nature, which was founded on strong theoretical and practical background.
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Critical review of the distribution of Equisetum × moorei and E. hyemale in the Nyírség (East Hungary)
131–144Views:456Confusing Moore's horsetail (Equisetum × moorei Newman) with rough horsetail (Equisetum hyemale L.) is a known problem in many countries. In our paper, we review some of the features used for their identification concerning their usability. We examined Equisetum hyemale stands in the Nyírség reported in the literature and E. hyemale specimens of the Herbarium of the University of Debrecen that were collected in the Nyírség. We found that most of these stands and herbarium specimens are actually E. × moorei. The occurrence of E. hyemale was only confirmed in Bátorliget. We also report several new occurrences of Equisetum × moorei in the Nyírség. To better separate the two taxa, we recommend micromorphological examination of the surface of the shoots, which can be performed in the field, as well as observing the height of the leaf-sheath.
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Floristic data from the northern edge of the floristic region ‘Crisicum’ (NE Hungary)
275-294Views:119This paper reports new floristic data of 161 taxa. Most of these data were collected between 2011–2013, and are coming from the northern edge of the floristic region 'Crisicum', the NE part of the Great Hungarian Plain, which is a little-known area from a floristic point of view. 43 CEU quarterquadrates and territory of 35 settlements are represented by the dataset. The paper reports occurrence localities for legally protected (e.g. Ophioglossum vulgatum, Stellaria palustris, Ranunculus polyphyllus, Thlaspi jankae, Lathyrus palustris, Elatine spp., Lycopsis arvensis, Lindernia procumbens, Cirsium brachycephalum, Cyperus pannonicus), strictly protected (Armoracia macrocarpa, Vicia biennis) and some little-known (e.g. Cardamine parviflora, Reseda luteola, Melilorus dentatus, Veronica catenata) taxa. Numerous data of alien species (e.g. Lepidium densiflorum, Euphorbia maculata, Sicyos angulatus, Erechtites hieracifolia, Elodea nuttallii, Elymus elongatus, Eleusine indica, Sorghum halepense, Eriochloa villosa) are also published. The most important data are documented by voucher specimens deposited in herbarium of Debrecen University (DE).
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Leucojum vernum at the Súri-Bakonyalja and overview of its distribution in the Bakony Region
242-246Views:347In Hungary Leucojum vernum is considered a plant species of montane character, distributed predominantly in Transdanubia. Apart from some isolated localities (Zalaszántó Basin Mts, Tapolca Basin), the occurrences of the species are mainly confined to the region “Magas Bakony” and its periphery within the Hungarian Middle Mts. In the spring of 2020 a large population was discovered by the author in the region “Súri-Bakonyalja”, which significantly expands the known local distribution range of the species. The new population, together with other phytogeographically significant species (e.g. Anemone nemorosa, Corydalis intermedia, Veratrum album), was found in a poorly explored valley system (Kisbér village: Báró-erdő). Based on all literature and herbarium records, the local distribution range of L. vernum is surveyed.
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The occurrence of Viola collina Besser in the Bükk and Uppony Mts. (N Hungary)
254-258Views:116Herbarium revisions and recent field observations revealed the occurrence of Viola collina Besser in the Bükk and Uppony Mts. (northern Hungary). This species has not been reported from the North Hungarian Mountains thus far. Altogether 94 localities of V. collina have been recorded in that region between 2007 and 2014. A map of the species’ local distribution is provided. Local habitat preferences of V. collina are briefly discussed.
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Contributions to the distribution of Arabis nemorensis (Cruciferae) in Hungary
155-169Views:94Arabis nemorensis is a poorly-known species with sparse and mostly uncertain literature records in Hungary. A complete list of its known occurrences was presented in Soó’s synopsis in 1968. A few new records of this taxon have been reported since then. In the present study all literature records were evaluated by means of revision of Hungarian herbarium materials. Several vouchers were succesfully traced, and their identity as A. nemorensis was confirmed (Baja, Ercsi, Lesenceistvánd, Miskolc, Nagykanizsa, Szigetújfalu). In other cases, the putative vouchers belonged to A. hirsuta or A. sagitatta, thus the corresponding literature records proved to be erroneous (Balatonmáriafürdő-alsó, Gyenesdiás, Kőszeg, Vasboldogasszony, Zalaújlak). On the other hand, hitherto unknown Hungarian localities of A. nemorensis were revealed, specifically in the Bükk Mts. (Eger, Kács), the Bakony Mts. (Márkó), in the western Balaton region (Hahót, Hévíz, Keszthely-Fenékpuszta, Nyirád, Tapolca) and the Hungarian Plains (Csákvár, Debrecen, Egeralja, Fertőd-Eszterháza, Kiskőrös, Lébény, Ócsa, Sárszentmihály, Szigetszentmiklós). The species appeared to be new for the phytogeographical regions Bakonyicum and Nyírségense. Additionally, a few specimens of A. nemorensis collected outside the territory of present-day Hungary were revised in BP. The historical occurrence of the species at Torja (Turia in Romania) is documented by two specimens (Schur, 1853, as A. gerardi; Jávorka & Keller, 1943, as A. hirsuta). Another specimen that was collected in 1879 at Óbecse (Bečej) and labelled as A. glastifolia by Kovács was found to be A. nemorensis too. Since the identity of Schneller’s literature record (1858) from Futak is uncertain, Kovács’s specimen is currently the only, though historical record of this species from Vojvodina province in Serbia. Further specimens of A. nemorensis were discovered in the collections of Lengyel. They were collected and labelled as A. sagittata near Pomogy (Pamhagen) in 1910 (and questionably in 1919), probably representing the first gathering of A. nemorensis from Burgenland state in Austria.
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Floristical data from abandoned vineyards of Cserhát Hills
260-266Views:42The results of floristic researches in abandoned vineyards of Cserhát Hills (Northeastern Hungary) between 2007 and 2014 are presented along with an account of relevant floristic literature and specimens of the Herbarium Carpato–Pannonicum Collection of the Hungarian Natural History Museum. Data of protected species and locally or generally rare ones are presented. Occurrences of Equisetum × moorei, Hesperis sylvestris, Gentiana cruciata, Scabiosa canescens, Campanula macrostachya, Carlina acaulis, Centaurea indurata, C. stenolepis, Epipactis atrorubens and Orchis ustulata subsp. aestivalis are especially significant. Some new localities of the following species are also emphasized: Pulsatilla grandis, Sorbus domestica, Centaurea scabiosa subsp. sadleriana, Orchis militaris, O. × hybrida, Ranunculus arvensis, Rapistrum perenne, Chamaecytisus virescens, Ch. austriacus, Hippocrepis comosa, Euphorbia salicifolia, Thymelaea passerina, Senecio erucifolius.
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Contributions to the flora of Western-Transdanubia (W Hungary)
213-234Views:105In this paper, we presented the results of our floristic survey in West-Hungarian region. The data were collected between 2005 and 2015 and supportted with herbaria or photo-documentation. This paper reports new floristic data of 98 taxa from West-Hungary with altogether 172 field-collected data records. In the survey we focused on little-known taxa with a phyto-geographic importance. For some taxa we checked the herbarium specimens of Savaria Museum and Natural History Museum of Budapest. During our fieldwork we recorded 8 species new to W-Hungary, including 3 species considered to be native to the region [Oenanthe banatica Heuff., Koeleria pyramidata (Lam.) P. Beauv., Polycnemum majus A. Braun.]. Almost all of the recorded species are regarded as indigenous plants, many of them has nature conservation importance, thus, their monitoring are highly advised. We found new localities of many species which of high importance for plant geography describing their current distribution. Most of the recorded species occur in natural or semi-natural habitats, but some taxa were found in synanthropic habitats and extensive arable lands.
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Fern (Pteridopsida) flora of South Nyírség (East Hungary)
162–182Views:342The study presents a number of new data on the fern flora of South Nyírség, and reviews all available literature and herbarium fern records from the area. A total of 19 Pteridopsida species were detected in the region. The hart's tongue fern (Asplenium scolopendrium) and the soft shield fern (Polystichum setiferum) were hitherto unknown in the whole Nyírség region (flora district “Nyírségense”). The broad buckler-fern (Dryopteris dilatata), scaly male fern (Dryopteris affinis) and floating fern (Salvinia natans) are new records to the area of South Nyírség. Dryopteris dilatata has been found in a surprisingly great number of localities. Some morphological characters facilitating identification of this species are also discussed. The local pine plantations, fens, carrs and humid deciduous forests are important habitats for ferns. In some localities the co-occurrence of 5-6 fern species were observed. The increasing aridity in the last decade is posing a serious threat to ferns in the area.
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Data to the flora of Heves–Borsod Plain I. Distribution of forest, forest steppe and steppe elements
16-65Views:201This paper reports new floristic data of 159 taxa (158 species and one hybrid) from the Heves–Borsod Plain (5 micro-regions between the Mátra and Bükk Mountains and the Tisza River valley). The data were collected between 1999 and 2018 and supplemented with collected specimen (altogether 93 herbarium sheets). This study is the first part of a thematic series that analyzes the distribution patterns of forest, forest steppe and dry grassland ‘steppe’ species, beyond reporting floristic data. The data were derived from 51 flora-mapping quadrats (CEU), based on 5.395 field collected data records. Some indicator species of the three species groups have been assessed in detail (with the addition of distribution maps of 25 species). I have also tested two East-West direction ‘lines’ (the 100-meter isoline and the Csörsz Ditch) for the possible existence of North-South chorological gradients in the area. True forest species (mostly Querco-Fagetea elements) show a dispersed pattern, their representatives can be linked to forest blocks appearing in the landscape. Streams from the adjacented hilly areas (even if they are strongly modified) are very important in the dispersal of the forest species. Their significance and role are decreasing towards the South. For some steppe and forest steppe species (e.g. Brachypodium pinnatum, Campanula bononiensis, Clematis recta, Elymus hispidus, Lychnis viscaria, Ranunculus illyricus, Sanguisorba minor, Stipa spp., Teucrium chamaedrys, Trifolium alpestre, Vinca herbacea) the examined lines indicate a regional area boundary, while for other species (e.g. Phlomis tuberosa, Thalictrum minus) gradient-like distribution differences were not observed. In the distribution of many species an additional southern gradient running along the northern boundary of the former Heves Floodplain was detected. Southwards to this line, the representatives of the selected species are already very sporadic, occurring only in synantropic habitats (e.g. Vincetoxicum hirundinaria).
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Flowering phenology and distribution pattern of Lizard Orchids (Himantoglossum)
157-167Views:82Hybridization is very rare between the species of Himatoglossum genus, despite the fact that it is common in other orchid genera. The reasons for this can be the lack of overlapping distribution areas or the different flowering period of the species. In the paper distribution and flowering data of Himantoglossum species are presented from herbarium sheets or references (sources paper-based or internet). We collected 1001 precisely dated flowering records of 12 species. In case of 6 species (H. adriaticum, H. caprinum, H. comperianum, H. hircinum, H. jankae, H. robertianum) we had sufficient records to calculate the average flowering day and blooming length. According to the collected data the distribution of the species are overlapping except for H. formosum, H. galilaeum and H. metlesicsianum. Studying 55 species pairs, insignificant differences in blooming time were only at 21 species pairs, among them only 5 (H. comperianum - H. caprinum, H. montis-tauri - H. comperianum, H. hircinum - H. adriaticum, H. montis-tauri - H. caprinum és H. jankae - H. calcaratum) have overlapping distribution area. Data on hybridization could be found only at three species pairs among them. From the results we can conclude that in the case of Himantoglossum genus the rarity of hybridization is mostly due to the phenological isolation.
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Lectotypification of Thesium kernerianum Simonk.
69-74Views:53Thesium kernerianum is an endemic species of the Eastern Carpathians (in broad sense). The paper includes a survey of the discovery as well as the literature and herbarium records of this species. The name is lectotypified, and a few inaccuracies appearing in the literature are corrected.