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Texture analysis as a method for grape berry characterization
Views:513The marketability of table grapes is mainly characterized by berry size, color, taste and texture. Mechanical measurements of table grape berries could provide objective information on the textural qualities of grape berries. In addition, this method might be suitable to study the effects of phytotechnical methods (such as girdling) on table grape quality. The aim of this preliminary work was to demonstrate how instrumental testing could be used to examine the effects of girdling on berry grape texture and define the textural characteristics of table grape berries. Cane girdling was carried out at veraison in two table grape varieties. Texture analysis was performed several times during the maturity. Besides this examination another five varieties were analyzed to assess their berry mechanical properties. Double compression test was used to determine berry hardness and its derived parameters. Puncture test was applied to evaluate skin hardness, skin elasticity and skin break energy. Skin thickness was also investigated. Berry hardness, skin hardness and skin thickness of the girdled grapevines were significantly affected by this technique. Most of the textural parameters showed differences among the seven cultivars.
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Microbial assessment of potential functional dairy products with added dried herbs
59-63Views:198The market of dairy products is a dynamically developing sector of the food industry. Potential, functional dairy products, made by adding herbs or spices, will have antimicrobial and antioxidant effect due to the active biochemical agents of the plant additives. Furthermore, these active components will widen the storage life of food products and enhance their organoleptic properties too. We worked out a technology for creating fresh cheeses using a gentle pasteurizing method by treating the mixture of raw milk and 1.5% fat contained in commercial milk. As herb additives, we used citronella (Melissa officinalis), and peppermint (Mentha x piperita) harvested by us and dried them via Tyndall-method in convective dryer on 40 °C for 5.5 hours per day. The drying period took three days. We bought dried citronella and mint from the supermarket, which were dried by ionizing radiation, to compare the microbiological pollution with the herbs dried by us.
The main target of this research was to create a microbiologically stable, potential functional dairy product. However, because of the bad quality of the raw milk and the gentle heat treatment we used for sterilizing bulk milk, or else, cheeses were not safe for human consumption. As a consequence, we need further studies to modify our technology and get a microbiologically stable product.
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The Effect of Zanthium Extracts on Germination in Relation to Interior and Exterior Factors
65-69Views:172In Hungary, the cocklebur species significantly endanger row crops (i.e. corn, sunflower, sugar beet). Their fast spreading is caused by many reasons: long-lasting emergence, reduced sensitivity to many kinds of herbicides, vigorous competitional ability, allelopathy, fast initial growth and changes in climate. The means of allelopathy is used by many species of plants in the competition with one another, as well as by the cockleburs. But the efficiency of this is influenced by many factors. Species with allelopathical effect could show different behaviour under different conditions: they can produce compounds with retarding and stimulating effects in different quantities or compositions. The actual condition of the acceptor plant and the mediator agent (i.e. soil) influence the final effect.
In this experiment influence of phenology and rainfall was studied on allelopathy of cockleburs in May, July and September. A stronger inhibition was observed on growth of cress before rain in July than in May, but this retarding effect disappeared after rainfall. However, the samples collected in September have significant inhibitory effect after rain. Inhibition of sugarbeet’s germination was stronger in May than in July, and the difference increased after rain. -
Possibilities of selection and production of blue and ornamental corn (Zea mays L.)
15-19Views:205The cultivation of alternative crops has an important role in world agriculture. Their market share is continuously growing in the food industry sector. In the present study, we show the cultivation and breeding perspectives of ornamental and blue corn. There exists possibility to cultivate ornamental corn in Hungary. There are many cultivars of ornamental corn, with various kernel colors, husks, stalks and leave colors. Blue corn is unique among other corns. Blue corn higher in protein, iron and zinc than commercial dent corn. Pests affective ornamental and blue corn are the same as those of other commercial corns. Ornamental and blue corns need to be grown a minimum of 500 m away from commercial yellow dent corn to minimize any cross pollination which may result in off-color kernel. Our aim in the presentation of this review was to broaden Hungarian literature.
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Effects of different groundcover matters on nutrient availability in an integrated apple orchard in Eastern-Hungary
21-25Views:284The aim of our study is to examine the effects of different groundcover methods on nutrient availability and uptake of apple orchard. The
experiment was carried out at the orchard of TEDEJ Rt. at Hajdúnánás-Tedej, in Eastern Hungary. The orchard was set up on lowland chernozem soil in the Nyírség region. It was established in the autumn of 1999, using Idared cultivar grafted on MM106 rootstocks at a spacing of 3.8 x 1.1 m.
The applied treatments were divided into two groups according to origins and effects. On the one hand, different livestock manures (cow,
horse and pig), on the other hand different mulch-matters (straw, pine bark mulch, black foil) were used. The different manures and mulches
were applied on the surface to test the effectiveness of these materials.
The effectiveness of manure treatments was higher than the other treatments on AL soluble P content of soil. Mostly the manure treatments
increased the AL soluble K of soil. Our all treatments increased 0.01 M CaCl2 soluble NO3 - -N content of the examined soil layers. The effect
of manure treatments was the highest. From the results it was evident that the amount of easily soluble organic nitrogen fraction distributed
more homogeneously than the other mineral N fractions examined.
Our results can be summarized as follows:
1. Our results pointed out that the used ground covering matters divided into several categories regarding its effect.
2. The available N, P and K contents of soil were mostly increased by applying manures.
3. The effectiveness of straw, mulch and mostly black foil was lower.
4. Differences were found between nutrient supplying treatments and the treatments which did not supply nutrients. -
Remembrance of Bognár Sándor (1921-2011)
16-20Views:164Dr. Sándor Bognár was a distinguished cultivator of the horticultural entomology and a determining personality of the higher education of Hungarian crop protection. Training of the scientific and extension specialists of Hungarian agricultural entomology would have been unimaginable without him. He was researcher of the Plant Protection Institute, innovator of the crop protection training’s essential and organisational standard at the Horticultural College and Faculty, who sacrificed his life’s work for developing the crop protection and for the tracking of the history of Hungarian crop protection. He dealt with the Pimplinae (Ichneuminidae) at the very beginning of his carrier and later with the difficulties of soil dwelling pests (Elateridae) but he dedicated a lot of time to the pests of rice and the pest-assemblages of fruit trees and grape. It is important to mention his activity on the phytophagous mites, thus one can call him as one of the founders of the Hungarian agricultural acarology. One top of his educational work was the wonderful manual „Agricultural Entomology” written with László Huzián in 1974 and 1979. László Szalay-Marzsó said of this book that it was impossible to write a better one. He has been right. In spite of his advanced age he participated systematically in the conferences in Hungary and shared the knowledge gained during his long life, and the morals and patriotism got from his models and developed considerably.
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Evaluation of various silo sorghum hybrids from the energetic aspect
107-111Views:226The sweet sorghum is a perspective plant of bioenergy, which can be the foundation of Hungarian bioethanol production in the future. By the examination six sweet sorghum hybrids have been examinated by the viewpoint of sugar aggregation and bioethanol production capacity. The founda tion of the surve y was three growing se ason (201 0, 2011, 2012 ). The ex per ime nt w as on the production sites of U niversity of Debrecen CAAES RISF Karcag Research Institute. The production site of the sweet sorghum have been sampled by samples of 1 m2 by hybrids within the period of august 15 and november 15. The sugar content of the samples have been measured by refractometer which was the base in the determination of ethanol production capacity.
As a continuation of the evaluation of energetic viewpoint the Higher Heating Value (HHV) have been measured from the bagasse of sweet sorghums. According to the results it can be stated that int he case of sugarcontent and the dinamics of the sugar aggregation can show several differences, while HHV of sorghum bagasse is within 16 200 and 16 900 J g-1.
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Seed treatment with Bacillus bacteria improves maize production: a narrative review
105-111Views:711Maize (Zea mays L.) is an important crop in relation to its production and consumption. Production of maize is constrained by soil infertility and poor quality seed. Microbial technologies like seed treatment with Bacillus bacteria improves the productivity of maize on infertile soil. However, due to variations in maize growth environments and Bacillus species, this review was conducted to identify the common species of Bacillus species used for seed treatment, and provide an overview of the effect of seed treatment with Bacillus on maize growth and yield. Results show that Bacillus subtilis, Bacillus pumilus and Bacillus amyloliquefaciens were the dominant species used for seed treatment. Bacillus was used as both a biofertiliser and biopesticide. The conspicuous positive effects of Bacillus were in plant height, shoot and root length, and shoot dry matter depending on the species. In terms of grain yield, Bacillus subtilis (8502 kg ha-1), Bacillus amyloliquefaciens (6822 kg ha-1) and Bacillus safensis (5562 kg ha-1) were the bacterial species that had an overall pronounced effect. The highest increase in grain yield was in the interactive effect of Bacillus megaterium + Bacillus licheniformis (18.1%) and sole Bacillus subtilis (15.6%), while Bacillus pumilus reduced grain yield by 4.8%. This shows that the improvement of maize productivity using Bacillus bacteria requires careful selection of the species for seed treatment.
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Use of molecular marker methods in the classification of bamboo taxa: A review
51-59Views:418Bamboo plants are currently attractive to researchers because of their versatile uses. Understanding the bamboos’ genetic level is needed to develop new varieties. Taxonomic identification is the basis for plant development. Bamboos were identified as their taxonomical morphological characters which are dependent on environmental factors. Molecular Marker techniques can be used to perform accurate genotype identification, which can be used for genetic diversity analyses. The RFLP, RAPD, AFLP, SSR, ISSR, iPBS, SCARS, SCoT, SRAP marker systems have been shown to be able to efficiently determine the genetic diversity of bamboo species based on genotyping. This paper summarizes research that aims to analyze the genetic diversity of bamboo species on a molecular basis.
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The influence of crop rotation and nutrition regime in winter wheat on the phytomass remains and the deposits of weed seeds remained in the ground
10-13Views:195An important problem refers to soil weeds seed bank with special reference to brown luvic soils and relationship between vegetal remains
(roots + stub) and nutrition regime – crop rotation plant. A great number of seeds can be found in first 20 centimeters of soil (plowing
layer). This fact causes a high weeds density, which has a negative effect on the cultivated plants leading to partial or total crop loss. -
Agricultural Biotechnology MSc. course and agriculture in the 21st century
208-214Views:194The establishment of an Agricultural Biotechnology MSc. course creates a new direction in Hungarian higher education. As an introduction, the article summarizes the main theoretical and practical possibilities and results of biotechnology which have necessitated launching the course. Subsequently, the preliminaries, aims and requirements of the Masters course are introduced. The main data of the agricultural MSc course at Szent István University in Gödöllő (Hungary) are the following: 4 semesters, 120 credits, 1200 contact hours, 4 weeks training in a biotech laboratory, 6 fundamental subjects, 8 basic subjects in biotechnology. Plant and animal biotechnology, are the two options for specialization and both have 10 separate professional subjects. The teaching staff consists of 34 teachers (93% of them have PhDs or higher scientific degrees) from 10 Departments of 3 Faculties of Szent István University.
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The effect of apoplastic pH on the nutrient uptake
65-71Views:184The pH of soil and rhizosphare –around the roots- determine the mobility and solubility of nutrients. The exudates organic acids of plant able to modify the pH, as well as the microorganisms also take part in mobilization of nutrients. The nutrient solve mostly in mildly acidic and neutral pH. The either assumption of utilization of nutrients is the uptake by roots and of course uptake to the cells to take part in metabolism. The pH of apoplast fluid determines the solubility and uptake of nutrients to the cells.
The aim of this study was to examine the effect of nutrient solution and apoplastic pH together with a bacteria based biofertiliser (Phylazonit MC®) on nutrient uptake and pH of apoplast fluid in case of nutrient solution grown plants in laboratory experiment. According to my results, the bicarbonate increased the pH of nutrient solution in due to influence the solubility and uptake of nutrients. The given bicarbonate to the nutrient solution and infiltrated into the apoplazma also modified the pH of the apoplast fluid of the test plants. The effect of bicarbonate and biofertilizer were different on the pH of the apoplast fluid and nutrient solution in nutrient solution experiment. -
Ideas on the European stone fruit yellows – as an entomologist can see them
30-34Views:300The European stone fruit yellows (ESFY) is an important endemic disease in Europe which causes in both, the Mediterranean countries and Central Europe serious damage. Its pathogen is the ‘Candidatus Phytoplasma prunorum’. The treatment and healing of the diseased trees and plantations with chemicals do not promise success. Thus, prevention may be the only solution. The transmission and spread of the pathogen happen by infected propagation material (grafting) or a vector (the psyllid, Cacopsylla pruni). Mechanism of the pathogen’s transmission and population dynamics of the vector have been extensively investigated in several European countries, which may allow by the control of C. pruni even to hold back the disease. Diseased stone fruit trees and wild Prunus spp. as main host species play an important role in maintaining and spreading the pathogen. C. pruni collects the pathogen by feeding on these plants and it carries persistently ‘Ca. P prunorum’. Researchers in Hungary have been characterized the disease only in terms of plant pathology, but neither the significance of the vector nor the role of wild Prunus spp. have been studied. This summary intends to give clues to these researches, that not only axe and saw should be the instruments of national control, but knowing the role and population dynamics of the vector the stone fruit production should be more successful.
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Effects of soil compost application on the leaf surface area per running meter of apple (Malus domesticaBorkh.) leaves cultivated in organic and integrated orchard
111-116Views:258The effect of different compost doses on the nutrient contents of the soil (in depth of 0–30) were measured in an (organic and integrated) apple orchards. The leaf surface area per running meter shoot for certain apple species (Golden Delicious and Pinova) were determined.
The field experiments were conducted in the Research Field of Debrecen-Pallag, University of Debrecen, the analysis of the soil and plant samples took place in the laboratories of the Institute of Agricultural Chemistry and Soil Science.
The compost treatments increased the amount of the examined nutrients in the top 30 cm of soil originated from organic and integrated orchard. Changes in rainfall were observed in the trends of the leaf surface area of the two apple varieties. 2010 was rainy and 2012 was drier. The leaf surface area of the Pinova increased better under the precipitation/year effect, than in case of the nutrient supply treatments. The opposite was observed in case of the Golden Delicious. -
Milestones in the development of agronomic management practices in crop production
203-209Views:245From the dawn of the history of the human race, agriculture has always been a profound activity of mankind producing food and feed as well as various plant originated materials for further processing.
Agronomy, like any other human activity, depends on the perpetual development of knowledge and technical skills, - in a modern context – science and innovation. This paper is intended to provide the reader with information regarding the main phases of the development of agricultural production from the Neolithic societies through the early Mesopotamian and Egyptian empires to the inventions of first organised learned society of Rome. The major research findings of the past two millennia including agro-chemistry, genetics and technical development are presented.
Such a review should not lead to any scientific conclusions, but rather a philosophical postulate similar to that of Jonathan Swift written some centuries ago: “And he gave it for his opinion, that whoever could make two ears of corn, or two blades of grass, to grow upon a spot of ground, where only one grew before, would deserve better of mankind, and do more essential service to his country, than the whole race of politicians put together”.
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The potential of biological control on invasive weed species
73-75Views:243Sorghum halepense is one of the invasive species in Europe. This study was made to identify the morphology of fungi on invasive weed species samples on the roots of Sorghum halepense. The samples were collected in the region of Debrecen. The experiment was conducted under laboratory conditions to determine the microscopic form of fungi. The samples were put on PDA and for identification of fungi is based on the morphological characteristics of the features and colonies of conidia that were developed in Petri dishes.
The examination of the culture revealed that the fungus from the root of Sorghum halepense was Aspergillus niger. Pathogenicity and the relationship between the fungus and Sorghum halepense are still uncertain so in the future pathogenicity tests and re-isolations from plants are very important steps.
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Global Issues of RangelandManagement
39-46Views:191Rangelands occupy about 50% of the world’s land area. They are ecologically and economically as important as rain forests and in even greater danger of degradation and disappearance. This paper reviews the definitions and distribution of rangelands and describes their global environmental importance in terms of erosion control, carbon storage and methane emission. Condition and degradation of rangelands are defined and discussed and it is argued that soil protection and carbon storage can be increased and methane emission per animal decreased by conservative use and improvement of rangelands, whilst at the same time alleviating hunger and malnutrition in developing countries. It is concluded that policies should be adopted by national governments and international deve-lopment programs to conserve and improve rangelands.
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The possibilities of biomass utilization in the field of spatial development
59-63Views:164By joining the European Union, Hungary made a commitment to increasingly utilize renewable energy sources. Keeping in view the agricultural circumstances of Hungary, we can state that biomass utilization has, overall, the greatest range of possibilities to use its products as ‘energy sources’. Biomass raw materials are useful in meeting emission control regulations for environment protection and to reduce climate change. The role of biomass production in spatial and rural development and in rural employment, and also in the decrease of Hungary’s energy dependence, supports development in biomass energy use. My aim in studying biomass utilization is the creation of a micro-regional level, decentralised, agro-energetic system-model for the use of plant and animal biomass for energy purposes, whether naturally or by anthropogenic activity, on agro- and sylvicultural areas, and on animal breeding farms. This model simulation method serves as a planning base for policy-makers during the spatial planning processes. The examination is presently in the stage of data collection. The collection of the basic data of the examination area and the purchase of the necessary equipment, materials and maps has already started. Contact has been made with local stakeholders, municipalities, municipality associations and micro-region managers.
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The impact of environmental factors on the measurement of the normalized difference vegetation index
141-147Views:358The level of nitrogen supply of a plant population can be quickly measured with non-destructive optical measurement devices and the differentiated determination of nitrogen shortage and the replenishment of nitrogen can also be carried out. The level of nitrogen supply is based on the fact that the chlorophyll content of crops is in close correlation with nitrogen content and that the amount of chlorophyll can be easily measured on the basis of the light absorption of chlorophyll molecules. The successfulness of optical measurements can be influenced by the change of weather parameters; therefore, it is important to know the correlations between measurement results and weather parameters when it comes to practical use.
The GreenSeeker Model 505 measurement device determines the relative chlorophyll content in the form of the Normalized Difference
Vegetation Index (NDVI) calculated on the basis of the intensity of the reflected red and infrared rays of light from the crop population. The measurements were performed in alfalfa population with 10 replications at five measurement heights and four measurement times. The weather parameters were measured by a weather station located in the middle of the alfalfa population and the correlations between the meteorological data and the NDVI values were examined. During the statistical evaluation of the results, it was established that the NDVI measurement is primarily influenced by the relative humidity of the air, secondly by air temperature and thirdly by wind speed. Relative humidity was in strong correlation with the NDVI values which were also influenced by the measurement height and time. Regression was not significant in the case of 20 cm measurement height, but the measurements above 40 cm height showed significant correlations. The correlation was shown to be strong at each measurement time, but the influence of humidity was the lowest at 11:00 and 14:00. -
Contexts between apple orchards with various cultivar comparisons and the effect of ATS (ammoniumthiosulphate) on fruit thinning
35-38Views:222Our research focuses on a fruit thinning material that can also be used in apple production. This material reduces significantly the required manual labor of hand thinning by russeting the blossoms selectively. The ATS (ammonium-thiosulphate) acts as chemical desiccant contrary to the nowadays commonly used materials such as naphthalene acetic acid, naphthyl acetamide, benziladenin and ethylene, which affect the metabolic processes of the plant by regulating the hormone system. In our experimentals cultivar ’Pinova’ and ’Golden Reinders’ were treated with different concentrations of ATS. The effect of these doses on the fruit setting and the quality and quantity parameters of the fruits was studied. According to our results, in the case of cultivar ’Pinova’ the ATS did not have any detected effects at the concentration of 1.5%. Application of ATS at 3% decreased considerably the fruit setting and fruit yield, accordingly the mean fruit size improved. The response to treatment in the case cultivar ‘Golden Reinders’ does not have any similar consistent results.
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Contributions to the 2014 and 2015 flight pattern and damages of Carpomyia schineri Loew.
96-99Views:193The fly Carpomyia schineri Loew is a pest of the berries of the feral Rosa canina group but is can also be found in the berries of grown rose cultivars. The larva damages the flesh of the berry, several larvae can develop inside a single berry. It can decrease the quantity and ascorbic acid content of the tea and jam, which are important human ascorbic acid sources during winter. At the same time in trapping experiments the adult can be confounded with adults of the walnut husk fly (Rhagoletis completa Cresson). In collections of berries at several sites in Hungary damage levels ranged from 0.88% to 65.08 %. Based on these damage levels we had the impression that the yellow sticky traps CSALOMON® PALz or PALs baited with the synthetic Rhagoletis lure caught moderate numbers of adults and were not satisfactory for detection, consequently improving trapping methods is necessary in the future.
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The strengthen of the second pillar in the European Union and Hungary
88-94Views:145The Common Agriculture Policy (CAP) took shape in the early 1960s. At that time the first action was to stop food shortages. Among the objectives of the Treaties of Rome appeared the subvention of rural development had not yet.
Rural development appeared, in 1992, in connection with accompanying measures, and by end of decade, the European Agricultural Model had taken shape. Agenda 2000 pointed out the direction of rural development and introduced the first and second pillars in the CAP. The regulations of rural development were simplified by 1257/1999/EC, which stressed the importance of rural development to and it has to continue the principle of subsidiarity and has to be decentralized.
The last enlargement raised new problems, the mid-term review of Agenda 2000 occurred and resulted in a new CAP-reform in 2003. The 1783/2003/EC rural development regulation modified the previous regulation. According to the new regulation it is necessary to strengthen the new rural development policy, enlarge the circles of accompanying measures, place greater emphasis on requirements of environment, human resource, animal welfare and plant health. Digression and modulation take part in the new CAP-reform in order to increase the role of rural development in the common budget.
In the next budget from 2007-2013, the European Union wants to create a single rural development fund and simplify programming, financing, monitoring. The EU wants to enlarge the instruments of rural development with a four axis model. -
The effect of increasing compost rates on the yield and nutrient content of ryegrass (Lolium perenne L.)
127-134Views:264Satisfaction of the increasing needs of humanity causes large environmental load. To provide a livable environment for future generations we have to satisfy our needs with the use of sustainable management. This is one of the biggest challenges of nowadays.
The amount of wastes emitted in increasing volume can be decreased by the recycling of them. The disposal of waste materials formed in the public spaces of cities and during the processing of agricultural row materials and by-products in landfills is inconceivable, so they must be recycled.
These materials mostly with organic compounds could be the primary substrates of composts. Completed with suitable additives, and applied appropriate treating technology, composts are capable to supply horticultural plants with nutrients. Composting wastes and byproducts not just decreases the amounts of deposited waste materials, but increases the nutrient (macro- and micronutrients) content of soils, so this is an environmentally friendly and alternative way of nutrient management of plants. -
The effect of different fertilizer treatments on the sulphur and protein content of wheat
73-76Views:360In this study the effect of N,P and K nutrients on the S and protein content of wheat grains was investigated in a long-term fertilization experiment set up in Nagyhörcsök. The calcareous chernozem soil having the following characteristics: pH (KCl): 7.3, CaCO3: 4.27%, humus: 3.45%, Al-soluble P2O5 and K2O: 60–80 and 180–200, KCl- Mg: 150–180, KCl+EDTA-soluble Mn-, Cu- and Zn-content: 80–150, 2–3 and 1–2 mg kg-1. The experiment had a split-split-plot design with 40 treatments in 4 replications. Plant samples were collected from 2002 and 2004. 2002 was a drought year while 2004 was very wet.
The main conclusions are as follows:
– The sulphur and protein content were than the control higher in every NPK treatments.
– The sulphur and protein content of the wheat grains were higher in 2004 that had a lot of rain than in 2002 that had drouht. -
Evaluation of sweet sorghum and sudangrass varieties by the viewpoint of bioethanol production
57-61Views:206Bioenergy and biofuels are very important in today’s energy policy. These kinds of energy resources have several advantages against fossil fuels. Environmental protection is a cardinal point of widespreading these technologies but the economic considerations are important as well. In order to improve the rate of the renewable energy in the energy consumption, the European Union settled down a program which determines a minimum ratio of renewable energy in the energy consumption for each member country of the EU. To fulfil the requirements bioenergy and biofuels should be produced. This production procedure needs adequate stocks which are commonly agricultural products.
One of the promising stocks is sorghum. This plant fits for bioethanol production due to its juice content being rich in sugar. In this study six sweet sorghum hybrids, two sudangrass hybrids and a sudangrass variety have been evaluated to determine their theorical ethanol production capacity.
On the score of the results of the year 2009 it can be set that sudangrasses have a lower theorical ethanol capacity than sweet sorghums have. In the case of sweet sorghums 1860.29–2615.47 l ha-1 ethanol yields had been calculated, while the sudangrasses had only 622.96–801.03 l ha-1. After that throughout three years (2011–2013) the sweet sorghum hybrids have been evaluated in order to determine the fluctuations of the ethanol production capacity caused by the impact of the years. As a result 2425.44–4043.6 l ha-1 theorical ethanol capacities have been calculated, which means that sweet sorghums can be an adequate stock to produce bioethanol.