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The herbarium of Debrecen University (DE) I. The “Rezső Soó Herbarium”
142-155Views:657The paper introduces the herbarium of Debrecen University (DE). All specimens of the herbarium were documented by digital photographs, and all data from the labels were entered into an Excel spreadsheet. The largest collection of Herbarium DE, the “Soó Rezső Herbarium” (abbreviation: SRH) consists of ca. 38.000 specimens from almost all European countries, but mainly from Hungary and the neighbouring countries. Most of the specimens originate from Hajdú-Bihar, Pest, Győr-Moson-Sopron, Borsod-Abaúj-Zemplén, Veszprém and Szabolcs-Szatmár-Bereg counties. The SRH stores a considerable amount of original/authentic specimens collected by Hungarian botanists, primarily by Rezső Soó and his followers, as well as Sándor Polgár. The most rapid growth of the SRH took place in the first half of the last century (1930–1950). By contrast, the collection was practically not developed between 1955 and 2000. After the millennium it started to grow again. Data on the most substantial part of the SRH covering native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting date, collector, file name of documentary photograph).
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The Herbarium of Debrecen University (DE) II.: The „Zoltán Siroki Herbarium”
15-22Views:924The paper introduces the “Zoltán Siroki Herbarium” (on the basis of the MS Excel spreadsheet database of the collection) which is a separate section within the Herbarium of Debrecen University (DE). The Zoltán Siroki Herbarium consists of ca. 20 000 specimens from almost all European countries, but mainly from the historical territory of Hungary. Although ca. 400 botanists contributed to this collection, most of the specimens were collected by Zoltán Siroki in Hajdú-Bihar, Pest and Borsod-Abaúj-Zemplén counties. The most rapid growth of the Herbarium took place in the middle of the last century (from the 1940s to 1960s), but Siroki remained an active collector until the 1980s. Some specimens from the Herbarium of former Royal Seed Testing Station (Budapest) was also incorporated in this collection; they represent gatherings from the last decades of the 19th century and first decades of the 20th century. Data on the most significant part of the collection, representing native plants collected in the present-day Hungary, are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).
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Herbarium database of the vascular collection of Eszterházy Károly College (EGR)
339-348Views:495The paper describes the vascular herbarium of the Eszterházy Károly College (EGR) in Eger (Hungary), according to its condition in 2013. All specimens of the herbarium were documented by digital photographs (ca. 8 000 specimens), and all data from the labels were entered into MS Excel spreadsheet. 54% of the specimens were collected in present-day Hungary, the other half comes mainly from the neighbouring countries, but more distant European countries are represented as well. Hungarian specimens were collected mostly in Heves, Pest, Zala, Vas, Borsod-Abaúj-Zemplén and Veszprém counties. Most of the herbarium sheets originated from the 1860’s and the 1950–60’s. Apart from these periods the collection has hardly developed. The most prolific collectors were Gábor Vida, Márton Vrabélyi, Árpád Károlyi and Tamás Pócs. More than half of the Hungarian flora (61%) is represented in EGR, although some taxa (e.g. Pterydophyta, Gymnospermatophyta) are much underrepresented. The digital photographs and the database are property of the authors and the Department of Botany and Ecology of Eszterházy Károly College. Data of non-cultivated plants collected in the territory of present-day Hungary are summarized in an electronic appendix (http://kitaibelia.unideb.hu/?download&aid=852&volume_id=94&lang=hun – including: catalogue number, taxon name, collector, settlement, date of gathering and file name of the documentary photograph). Further data can be required from the corresponding author or the curator of the herbarium.
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In memoriam Zoltán Siroki (1906–1987)
3-14Views:278Zoltán Siroki (1906–1987) was a Hungarian agronomist and lecturer. His professional activity was connected mainly to Debrecen. Beside his widely known ornithological studies his floristic and fitocoenological data to Matricum, Crisicum and Nyírségense Hungarian regions are also important. Herbarium of Zoltán Siroki, consisting of ca. 20 000 specimens is a remarkable part of the Herbarium of Debrecen University (DE).
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The historical occurrence of Digitaria ciliaris in Hungary – results of a herbarium survey
53–62Views:412Revision of genus Digitaria in BP Herbarium Carpato-Pannonicum collection helped to discover the historical occurrence of Digitaria ciliaris (Retz.) Koeler from Győr, in the area of an old cotton and wool processing and textile factory. However, it has been proved that the previously published Hungarian records of the species are all wrong, as the Digitaria-keys of Hungarian herbals misinterpreted the taxon, mainly as a synonym of D. sanguinalis subsp. pectiniformis. The herbarium survey confirmed that D. sanguinalis is the most common taxon in Hungary. D. sanguinalis subsp. pectiniformis is also widespread, but probably somewhat under-mapped. Probably not native to Hungary, most probably an archeophyte taxon. Specimens have been collected from disturbed habitats associated to railways and from xerothermic vineyard areas for over a hundred years. D. ischaemum is certainly native to the region, with a recognisable natural distribution pattern, and is found mainly in the sandy and gravelly soils of the lowland and hilly areas of the country. This species has recently become more common in urban areas across the country with the recent spread of sand-lined pavements. D. ciliaris of tropical/subtropical origin is a casual alien species that was formerly present in the Hungarian flora.
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The herbarium of the Botanical Garden of Eötvös Loránd University (BPU)
55–59Views:673The paper introduces the herbarium of Eötvös Loránd University (BPU), currently stored at the Botanical Garden of the University, according to its state in the year 2013. The BPU herbarium consists of ca. 16 000 specimens the relevant data of which have been organised into an OpenOffice Calc spreadsheet database. The specimens were collected in 16 European countries, mainly in the current territory of Hungary (80%), Romania (12%) and Slovakia (5%). Other countries are represented with very few specimens. The vast majority (ca. 70%) of the Hungarian specimens were collected in Pest, Veszprém, Borsod-Abaúj-Zemplén, Hajdú-Bihar and Szabolcs-Szatmár-Bereg counties. Although the gatherings cover a century and a half period of time, the most rapid growth of the collection took place from the 1930s to the 1960s. The most prolific collectors were Rezső Soó and his followers: Lajos Felföldy, Tibor Simon and Szaniszló Priszter. Felföldy and Simon enriched the collection through more than 60 years. Data on the native plants collected in the present-day Hungary are summarized in Electronic Appendix 1. (incl. taxon name, settlement, collecting year, collector, file name of documentary photograph).
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Data on the occurrence of sedge species (Carex, Cyperaceae) in and around South Nyírség (East Hungary)
165-198Views:599We provide floristic data on a total of 38 sedge species in the area of South Nyírség and its immediate surroundings. We also detected some populations of previously undescribed or rare species in the Hungarian Great Plain, such as Carex buxbaumii, Carex hartmanii, Carex umbrosa, and Carex paniculata. We report several new occurrences of species with little literature data from the area, such as Carex brizoides, Carex michelii, Carex pallescens, Carex elongata. However, during the last 13 years, we were not able to detect seven species previously mentioned in the literature. Herbarium data confirmed that some of the older Carex nigra occurrences for the area actually refer to Carex cespitosa. Based on herbarium data and the literature, we think that local data on Carex pairaei may largely refer to Carex divulsa subsp. leersii and/or Carex spicata. Contrary to many pieces of literature, Carex vulpina was not observed inside the sandy area.
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The ‘rediscovery’ of the Birdsfoot Fenugreek (Trifolium ornithopodioides (L.) SM.) in the Hortobágy, Hungary
207–212Views:528In 2009, the first author discovered an unknown population of Trifolium ornithopodioides in the Ágota-puszta of Hortobágy plain. In 2016, the second author found an another population in the vicinity of Karcag (Ecse-zug puszta). Up to now there was only one published occurrence of this species from the region. The specimen collected by Szujkó-Lacza Júlia, Kováts Dezső and Fekete Gábor in 1974 was deposited in the herbarium of the Hungarian Natural History Museum (BP), but this incomplete specimen was misidentified, and not Trifolium ornithopodioides. Therefore, this newly discovered site at Ágota-puszta should be regarded as the first trustworthy occurrence of this species at the Hortobágy plain.
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On the former occurrence of Calla palustris in Hungary
200-210Views:1974The origin of a more than 150 years old herbarium specimen of Calla palustris L. stored in JPU is reconsidered in this study. According to the information on its label, the specimen was collected in 1856 at Lake Velence, close to Székesfehérvár city (central Hungary). Some Hungarian botanists are skeptical about the authenticity of this record. Based on the current occurrences of some remarkable swamp species (e.g. Carex pseudocyperus, Liparis loeselii, Sphagnum spp.) and habitat types (e.g. willow carrs and swamp forests) at the lake, as well as the habitat preference of bog arum, there is no reason to question the former occurrence of the species here or even in other Hungarian localities. The content and accuracy of the examined voucher meet the expectations of floristic data. However, in the absence of recent data the species can be considered extinct from Hungary.
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Dr. Sándor Polgár was born 140 years ago
169–184Views:399Dr. Sándor Polgár was the most outstanding botanist of Győr county (NW Hungary). He provided important results in floristic studies, phytogeography, taxonomy and in the research of alien plants. „Győr megye flórája” („Flora of Győr county”), published in 1941, was one of the most important monography in his period. He is the author of the rare, hybridogenous species Ornithogalum ×degenianum, known only from Hungary. His private herbarium was one of the biggest in Hungary with more than 20,000 specimens. As a teacher he taught geography and nature studies in his home town Győr between 1900 and 1935. Because of his Jewish origin, he was a victim of the holocaust in 1944.
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Dr. Antal Waisbecker was born 180 years ago
179–192Views:277Dr. Antal Waisbecker (1835–1916) was a Hungarian physician and the most prominent botanist of Vas county (West Hungary) at the turn of the 19–20th century. He contributed with important floristic data for the botanical monography of Vas county, authored by Vince Borbás. He processed the pteridophytes of Vas county and cormophytes in the surroundings of Kőszeg. Antal Waisbecker was an expert in the knowledge of sedges too. He discovered and described the Carex fritschii. Remarkable amount of herbarium specimens collected by Waisbecker are deposited at the Savaria Museum (Szombathely, SAMU) and Hungarian Natural History Museum (Budapest, BP).
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Contributions to the lichen flora of the Mecsek Mts (South Hungary)
16–26Views:253A checklist of 115 lichen taxa was compiled from the Mecsek Mts, based on floristical records of 221 old or recently collected herbarium specimens (incl. 37 duplicates) deposited in the JPU herbarium (Pécs, Hungary). The occurrence of 22 taxa are new for the Mecsek Mts: Arthopyrenia cinereopruinosa, Baeomyces rufus, Candelariella reflexa, Catapyrenium rufescens, Cladonia macilenta, Dimerella pineti, Hypogymnia tubulosa, Ochrolechia arborea, Pleurosticta acetabulum, Pseudevernia furfuracea, Psilolechia lucida, Pycnothelia papillaria, Rinodina sophodes, Tomasellia arthonioides, Toninia candida, Trapelia involuta, Varicellaria hemisphaerica, Verrucaria marmorea, V. nigrescens, V. parmigera, Xanthoparmelia conspersa, X. verruculifera. One legally protected lichen species (Peltigera leucophlebia) is also represented by two specimens.
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Lajos Felföldy: a prominent Hungarian botanist and hydrobiologist
3–25Views:606Lajos Felföldy (1920–2016) was one of the most versatile and open-minded Hungarian biologists. He began his scientific career as a student of Prof. Rezső Soó. Between 1938 and 1946 he participated in geobotanical studies in University of Debrecen and Kolozsvár (Cluj-Napoca, Romania). He contributed with important achievements to the development of several biological disciplines. His pioneering study on the effects of air pollution on epiphytic lichens (1942) was among the firsts in the world. He described Hemitherophyte life-form as a discrete unit within Raunkiaer's plant life-form system (1942). He was a pioneer in the cytological (caryological) study of wild vascular plant species in Hungary (1947–1949). His results regarding to primary production of freshwater algae and algal culture (1958–1960) were in leading edge. In 1972 he founded and until 1990 edited the Hungarian series entitled ‘Vízűgyi Hidrobiológia’. Books of this series aimed to publish identification keys of freshwater taxa. These books were proved to be decisive and useful tools for Hungarian hydrobiologists in biological classification of brooks, streams, rivers and different types of stagnant waters. Between 1934 and 2009 he collected more than ten thousand herbarium sheets. After his retirement, he dealt with the revision of the herbarium material of Department of Botany in Hungarian Natural History Museum and Botanical Garden of Eötvös University (Budapest). He participated in the preparation of the New Hungarian Herbal. His scientific work was characterised by deep humility for nature, which was founded on strong theoretical and practical background.
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Critical review of the distribution of Equisetum × moorei and E. hyemale in the Nyírség (East Hungary)
131–144Views:678Confusing Moore's horsetail (Equisetum × moorei Newman) with rough horsetail (Equisetum hyemale L.) is a known problem in many countries. In our paper, we review some of the features used for their identification concerning their usability. We examined Equisetum hyemale stands in the Nyírség reported in the literature and E. hyemale specimens of the Herbarium of the University of Debrecen that were collected in the Nyírség. We found that most of these stands and herbarium specimens are actually E. × moorei. The occurrence of E. hyemale was only confirmed in Bátorliget. We also report several new occurrences of Equisetum × moorei in the Nyírség. To better separate the two taxa, we recommend micromorphological examination of the surface of the shoots, which can be performed in the field, as well as observing the height of the leaf-sheath.
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Sándor Polgár and the research of the adventive flora of Hungary
188–197Views:365In the 20th century Sándor Polgár was one of the most outstanding researchers of adventive plant species in Hungary. He wrote nine publications in this theme. In the industrial environment of his home town Győr (NW Hungary) he found tropical species, most of them were southern-American origin. The richest alien flora came around the oil factories of the town. He reported 65 taxa new for the Hungarian flora, four of them were new for Europe too. He was a great expert of problematic Solanum, Amaranthus and Chenopodium genera.
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Buglossoides rochelii (Boraginaceae) – a supplement to the flora of the Pannonian Plain
187–192Views:222Previously unknown occurrences of Buglossoides rochelii (Friv.) Stoyanov, Mátis & Sennikov (≡ Lithospermum rochelii Friv.) have been identified in the Pannonian Plain, based on field investigations and a critical revision of the genus Buglossoides in the major herbarium collections of Hungary and Serbia. In both its eastern range and the Pannonian Basin, the species predominantly occurs as a psammophyte. This paper summarizes its historical distribution in Hungary and Serbia, and provides new data on the current status of its habitats and populations in Serbia.
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Distribution and identification key of Rubus L. ser. Micantes in Hungary
56-68Views:163Based on a substantial field-work and herbarium revision carried out between 2007 and 2014, the authors present the taxonomical evaluation of recently known representatives of Rubus L. ser. Micantes Sudre in Hungary, providing the first grid-based distribution maps and an identification key of four species of the section (Rubus balatonicus Borbás, R. clusii Borbás, R. styriacus Halácsy, R. tabanimontanus Figert) for Hungary.
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Contributions to the flora of the Heves–Borsod and Uppony Hills and adjacent territories
173–226Views:292Results of nearly two decades of floristic research in the territory of Heves–Borsod Hills (Tarna-vidék) and the northern foreground of the Bükk Mts are presented in this paper (altogether 4421 records; BÁ: 1665, SJ: 2752). Several old literature records – partly supported by vouchers – are confirmed, e.g. Hypericum elegans and Lappula heteracantha from the Uppony Gorge, as well as Cephalaria transsylvanica, Cypripedium calceolus, Diplotaxis erucoides and Plantago indica from the Heves–Borsod Hills and the northern foreground of the Bükk Mts. Other former literature records (partly from the authors) are revised. These revisions are mainly due to changes in taxonomic concepts of some genera in new keys (Carex, Chamaecytisus, Epipactis, Molinia, Sorbus). Old literature and herbarium records of other significant taxa (Onosma visanii, Scutellaria columnae, Sorbus sp., Utricularia bremii/minor) are revised too. The new occurrence of Ferula sadleriana in the Uppony Gorge is probably the result of intentional seed dispersal. Of the many rare plants listed in our paper some are new for the flora of the region under study: Agrimonia procera, Alchemilla micans, Asplenium adiantum-nigrum, Astragalus austriacus, A. exscapus, Blysmus compressus, Carex appropinquata, C. cespitosa, Catabrosa aquatica, Centarea indurata, Dactylorhiza × aschersoniana, Epipactis voethii, Epipogium aphyllum, Equisetum hyemale, Festuca drymeja, Gagea bohemica, Glyceria nemoralis, Hesperis sylvestris, Myosotis caespitosa, Phegopteris connectilis, Platanthera chlorantha, P. × hybrida, Rosa gizellae, Scilla kladnii, Senecio doria, Solanum villosum, Taraxacum palustre, Triglochin palustre. Two phytocoenological relevés representing the habitats of Myosotis caespitosa and Spiraea media are presented. Currently known local distribution of some montane and forest-steppe elements are shown on maps.
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In memoriam János Hulják (1884–1942)
5-10Views:286János Hulják prominent Hungarian botanist was born 130 years ago. He worked as teacher and director in elementary school at Pereces (today it belong to Miskolc). He played key role in the floristic exploration of Northern Hungarian Mountain Range. His huge herbarium collection was destroyed during World War II., only the exchange material was preserved (about few hundred sheets), mainly in BP and DE. Some vascular plant species [Centaurea huljakii J. Wágner, Sorbus huljakii Kárpáti, Galium × huljakii Jávorka and Pilosella auriculoides subsp. huljakii (Zahn) Soják] were named after him.
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The clammy goosefoot (Chenopodium pumilio R.BR.) in Zugló (Budapest) and new data on the distribution of invasive species in NE Hungary
221-226Views:387The paper summarise occurrences of some rare invasive weeds of Hungary. We found currently second population at the Australian origin Chenopodium pumilio in Budapest (Zugló) in September 2016. Further, 27 occurrence data of 8 other species (Amaranthus deflexus, Echium maculatum, Lepidium densiflorum, Oxybaphus nyctagineus, Panicum dichotomiflorum, Portulaca grandiflora, Sarothamnus scoparius, Tragus racemosus) is also reported in the persent paper from the ‘Északi-középhegyég’ area (Northeast-Hungary), and some data from beside the Hungarian–Slovakian border, especially settlements, strongly disturbed places (especially road curbs, pavement cracks, train stations), indicating the Central European flora mapping system quadrate number. Voucher specimens were deposited in herbarium of Hungarian Natural History Museum, Budapest (BP).
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Taxonomic revision of alien Ludwigia species in Hungary
153–164Views:614Some tropical Ludwigia species have been planted as ornamentals in Hungarian thermal springs since the 1920s. Although they have become naturalised or even invasive in a few places, their specific identity remained generally uncertain. Our herbarium and field studies revealed that three species of alien water-primroses occur in Hungary. The most frequently planted one is Ludwigia repens, which is, however, a sporadic species in its native distribution area. We have found this species in several parts of the country. Although a few escaped and self-sustaining populations were also observed, this species has appeared to be non-invasive so far. Ludwigia grandiflora appeared in Hungary at the beginning of the 2000s. At the moment it is known from three localities, though spreading rapidly, becoming already invasive in its locality at Tata city. A similar species, L. peploides has recently been reported from two localities, where it forms small stands; the success of its naturalisation can be assessed in the future. We found no evidence for the Hungarian occurrence of L. alterniflora that had been published from Eger city. Therefore we suggest deleting this species from the current Hungarian checklist.
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Data on alien vascular plant species in Hungary I. (1–6)
65–80Views:912This is the first in a series of papers by various authors presenting previously unpublished data relevant to the knowledge of alien vascular plants in Hungary. This paper deals with the families Pteridaceae, Brassicaceae, Plantaginaceae, Rosaceae, Ranunculaceae, Fabaceae and Poaceae. It includes new area records and distributional considerations for Cochlearia danica and Plantago coronopus spreading along Hungarian road-network as well as records of Panicum riparium from the Northern Hungarian Mountain range based on field work and herbarium revision. We note the importance of checking the data previously collected as P. capillare. Floristic data of two species not previously recorded in Hungary (Cardamine corymbosa, Ranunculus marginatus) and 3 sporadically occurring alien species (Aphanes australis, Lepidium didymum, Medicago arabica), from garden centres in the Debrecen area were published. A 7-year long survival of a population of Adiantum capillus-veneris in a traditional, abandoned well is also documented between 2017 and 2024. The paper reports that traffic infrastructure can facilitate the spread of Elymus elongatus along roads.
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Contribution to the distribution of Epipactis species in the Keszthely Mts
8-14Views:217In this paper I present occurrence data of Epipactis species in the Keszthely Mts which were collected during the last four years. Ten species had literary or herbarium data from the area previously, all of them has been confirmed (Epipactis atrorubens, E. helleborine, E. leptochila, E. microphylla, E. moravica, E. muelleri, E. palustris, E. purpurata, E. nordeniorum, E. voethii) and another five species (Epipactis albensis, E. neglecta, E. peitzii, E. pontica, E. tallosii) furthermore a hybrid (Epipactis ×reinekei) were found newly on the area.
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Historical occurrence of Spergularia echinosperma, a rare Nanocyperion species in Hungary
3–9Views:487Three species of the genus Spergularia have been known from Hungary. The most widespread one, Spergularia rubra (L.) J. Presl et C. Presl occurs on young sandy fallow fields and other disturbed habitats. The other species, S. marina (L.) Bess. and S. media (L.) C. Presl, occur sporadically in saline grasslands, and both have been recently spreading along salted roads. During a revision of the genus Spergularia in Hungarian herbaria, specimens of the species S. echinosperma (Čelak.) Asch. et Graebn. were found. It is a rare species previously known to occur only in Czechia and Germany and to be extinct in Austria and Slovakia (a few occurrences along the Morava and Danube rivers between 1909 and 1953). However, it was recently discovered in Western Siberia and Kazakhstan. It is a typical mudflat plant that occurs on exposed sandy shores of water bodies. Two herbarium sheets collected by Gyula Tauscher from near the village of Ercsi (riverbank of the Danube) in 1903 were found in the Carpato-Pannonicum collection of the Hungarian Natural History Museum. In addition, a new specimen was found, collected in Slovakia (Danube near Šamorín, collected by Mihály Résely in 1868) at the vicinity of the Hungarian border. The species was not confirmed during a field inspection of the target localities in Hungary; therefore, we have to consider it extinct in the country as well as in the whole Pannonian Basin. Recently, S. echinosperma was split into two subspecies – subsp. echinosperma and subsp. albensis. The subsp. echinosperma is probably restricted to fishponds in the SW part of Czechia and subsp. albensis occurs mainly in the floodplains of the river Elbe in Germany. The extinct populations of S. echinosperma in the Pannonian Basin belonged to the subsp. albensis. This taxon is morphologically close to S. rubra but can be reliably distinguished by densely verrucose seeds and shorter than wide stipules.